1. Family: Lamiaceae Martinov
    1. Genus: Pycnostachys Hook.
      1. Pycnostachys niamniamensis Gürke

        This species is accepted, and its native range is South Sudan to Kenya.

    [FTEA]

    Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Pycnostachys, G Bramleyshaped. Flora of Tropical East Africa. 2009

    Type
    Type: Sudan, Niamniam, east of R. Huuh, Schweinfurth 3750a (B†, holo. UPS, iso., K!, fragment)
    Habit
    Perennial herb 0.5–1 m tall, becoming woody below
    Stem
    Stems erect, slightly branching, quadrangular, or rounded-quadrangular where woodier, pubescent with short eglandular, closely appressed hairs, also with red sessile glands
    Leaves
    Leaves opposite or ternate, sessile or minutely petiolate; blades elliptic to narrowly elliptic, 2–10 × 0.8–5 cm, margins subentire to serrate or crenate-serrate in the upper half, apex acute to obtuse, base sessile or obtuse where minutely petiolate, subglabrous or pubescent with scattered short eglandular hairs, these more plentiful on the venation which is sunken above and raised beneath, also with yellow sessile glands above, red sessile glands beneath; petiole 1–2 mm long
    Inflorescences
    Inflorescence dense, when in flower subglobose, in fruit 25–75 × 18–28 mm; bracts subtending entire inflorescence lanceolate, 6–15 mm long, acuminate, enclosing buds then becoming deflexed in flower, pubescent with short eglandular hairs, persistent through flowering stage; bracts subtending single flowers sometimes forming a coma at the tip of the inflorescence, lanceolate with a short to long acuminate apex that sometimes curls over, 4–8 mm long, pubescent with short eglandular hairs, ciliate
    Calyx
    Calyx with tube 0.5–1 mm long at anthesis, outer surface with eglandular hairs and yellow and red sessile glands, inner surface with ± 0.5 mm long narrow triangular scales present between calyx lobes; lobes 5, subulate, slightly curved, with a membranous wing either side of the midrib, ± subequal, 4–6 mm long, pubescent with short eglandular hairs or ± glabrous; fruiting calyx with tube ± 4 mm long, somewhat compressed dorsally, ventrally gibbous, lobes curved, apparently not lengthening
    Corolla
    Corolla white or pale blue, 10–11 mm long; tube pubescent with eglandular hairs, also with yellow sessile glands, very narrow and parallel-sided for ± 3 mm, then curving sharply downwards and broadening for ± 3 mm before opening out into two lips; posterior lip shorter than anterior, 2.5–3 mm long, with yellow sessile glands, densely pubescent with eglandular hairs; anterior lip ± 4 mm long, 2–3 mm deep, the front densely pubescent with eglandular hairs and yellow sessile glands
    Stamens
    Stamens connate for ± 1.5 mm, barely exserted
    Fruits
    Nutlets (immature) ± 1 mm long.
    Ecology
    Swampy grassland; 900–2100 m
    Conservation
    Known over a considerable area in Kenya and Uganda but habitat under some threat from agriculture; possibly threatened.
    Note
    There is some variation in leaf indumentum and calyx lobe length – specimens from Uganda are ± glabrous, but those from Kenya tend to have very short hair on the lower leaf surface, especially on the venation. Sudanese collections can have calyx lobes up to 10 mm long. P. niamniamensis is very closely related to the more southerly distributed P. prittwitzii, but can be distinguished from it because of its elliptic or narrowly elliptic rather than oblanceolate leaves, usually less conspicuous bracts, and longer calyx lobes (4–6 mm not 3–4 mm). It is also closely related to P. reticulata and could be confused vegetatively with the Tanzanian collections, but can be distinguished by its dorsally compressed fruiting calyx, which has longer, curved lobes, and its wider fruiting inflorescence. It also apparently has more northern distribution than P. reticulata, only reaching as far south as K 6.
    Distribution
    Range: Sudan and possibly Ethiopia Flora districts: U1 K2 K3 K5 K6

    Images

    Distribution

    Native to:

    Kenya, Sudan, Uganda

    Pycnostachys niamniamensis Gürke appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1914 Scott Elliot, G.F. [7155], Kenya K000405971 Unknown type material
    Jan 1, 1914 Petherick [s.n.] K000405959 Unknown type material
    Schweinfurth [3750a], Sudan K000405960 Unknown type material
    Lindblom [s.n.], Kenya K000405972 Unknown type material

    First published in Bot. Jahrb. Syst. 22: 145 (1895)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Paton, A.J., Bramley, G., Ryding, O., Polhill, R., Harvey, Y., Iwarsson, M., Willis, F., Phillipson, P., Balkwill, K., Lukhoba, C., Otiend, D & Harley (2009). Lamiaceae (Labiatae) Flora of Tropical East Africa: 1-430.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora of Tropical East Africa
    • U.K.W.F. ed. 2: 290, t. 128 (1994), pro majore parte.
    • F.P.S. 3: 223 (1956)
    • K.B. 1939: 584 (1940)
    • N.B.G.B. 8: 71 (1921)
    • F.T.A. 5: 383 (1900)
    • E.J. 22: 145 (1895)

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0