Skip to main content
  1. Family: Lamiaceae Martinov
    1. Genus: Clinopodium L.
      1. Clinopodium vanum (Epling) Harley & A.Granda

        This species is accepted, and its native range is S. Peru to W. Bolivia.


    Wood, J.R.I. Kew Bull (2011) 66: 199.

    Much-branched, erect, aromatic undershrub 0.5 – 1 m high, the ultimate branches slender, straight to slightly bent; young stems quadrangular, puberulent apart from the pilose leaf nodes
    Leaves petiolate, not crowded towards the branch tips; petioles 1 – 1.25 mm long; lamina 4 – 22 × 2 – 16 cm, broadly-lanceolate, ovate or ovate-elliptic, acute, basally cuneate, obscurely toothed, minutely puberulent, paler and gland-dotted beneath
    Inflorescence of 1 – several, axillary cymes, these apparently always 3-flowered but the two lateral flowers often not developing fully so apparently 1-flowered; peduncles c. 0.75 mm, pubescent; bracts 0.75 – 1 mm long, subulate, pubescent; pedicels c. 0.75 mm, pubescent; calyx 2 – 3 mm long, cylindrical, the exterior ribbed, pubescent, gland-dotted, the interior pilose at the mouth, teeth c. 1 × 0.75 mm, narrowly triangular; corolla white, pubescent and gland-dotted on the exterior, infundibuliform, the tube 5 – 8 mm long, weakly 2-lipped and subequally 5-lobed, the lobes 1.25 × 1.25 mm, rounded; stamens didynamous, filaments glabrous, the shorter pair c. 1.5 mm long, the longer pair c. 2 mm long, anthers at mouth or weakly exserted; style c. 6 mm long, glabrous
    Nutlets c. 1.5 × 1 mm, trigonous, somewhat flattened, obtuse, glabrous.
    A rather local species of dry, high Andean slopes in puna vegetation between 3100 – 4000 m in southern Peru and northern Bolivia. Map 3.
    Least Concern (LC).
    Clinopodium vanum appears to flower throughout the year including the summer rainy season from November to April when C. bolivianum is not usually flowering.
    A few specimens cited above (Beck 4112, 4226, 11356, Boeke 2992, Wood & Goyder 15355) show a degree of introgression with Clinopodium bolivianum and are quite possibly of hybrid origin as typical C. vanum and C. bolivianum occur in the area where they were found, although this is only inference. These intermediate plants have glabrous nutlets as in C. vanum but the apex is acuminate, the leaves are oblong-elliptic rather than ovate, and the axillary cymes are few-flowered, often with only a single flower (Fig. 9). They seem always to be fertile but, as in the plant illustrated, the stamens may be somewhat dimorphic on the same plant.



    Native to:

    Bolivia, Peru


    Other Data

    Clinopodium vanum (Epling) Harley & A.Granda appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Feb 23, 2017 Arroyo, L. [3874], Bolivia K000543876
    Weberbauer, A. [7259], Peru K000193636 Unknown type material


    First published in Kew Bull. 55: 926 (2000)

    Accepted by

    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741.
    • Wood, J.R.I. (2011). Clinopodium L. (Lamiaceae) in Bolivia Kew Bulletin 66: 199-226.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.


    Kew Bulletin

    • Schmidt-Lebuhn, A. N. (2008). A Revision of the Genus Minthostachys (Labiatae). Mem. New York Bot. Gard. 98: 1 – 75.Google Scholar
    • Hughes, C. E. & Eastwood, R. J. (2006). Island Radiation on a continental scale: exceptional rates of plant diversification after uplift of the Andes. Proc. Nat. Acad. Sci. 103: 10334 – 10339.PubMedCrossRefGoogle Scholar
    • Bell, C. D. & Donoghue, M. J. (2005). Phylogeny and biogeography of Valerianaceae (Dipsacales) with special reference to the South American valerians. Organ. Divers. Evol. 5: 147 – 159.CrossRefGoogle Scholar
    • IUCN (2001). IUCN Red List Categories and Criteria: Version 3.1. Prepared by the IUCN Species Survival Commission. IUCN, Gland, Switzerland & Cambridge, UK.Google Scholar
    • (2000: 926).
    • Harley, R. M. & Granda Paucar, A. (2000). List of species of tropical American Clinopodium (Labiatae) with new combinations. Kew Bull. 55: 917 – 927.CrossRefGoogle Scholar
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1). Continental Publishing, Deurne.Google Scholar
    • Xifreda, C. C. (1999). Lamiaceae. In: O. Zuloaga & O. Morrone (eds), Catálogo de las Plantas Vasculares de la Republica Argentina II, Angiospermae (Dicotyledoneae). Monogr. Syst. Bot. Missouri Bot. Gard. 74: 769 – 781.Google Scholar
    • Cantino, P. D. & Wagstaff, S. J. (1998). A Reexamination of North American Satureja s.l. (Lamiaceae) in light of molecular evidence. Brittonia 50: 63 – 70.CrossRefGoogle Scholar
    • Wagstaff, S. J., Olmstead, R. G. & Cantino, P. D. (1995). Parsimony analysis of cpDNA restriction site variation in subfamily Nepetoideae (Labiatae). Amer. J. Bot. 82: 886 – 892.CrossRefGoogle Scholar
    • Torrico, G., Peca, C., Beck, S. G. & García, E. (1994). Leñosas Utiles de Potosí. Potosí.Google Scholar
    • Pontiroli, A. (1993). Labiatae. In: A. L. Cabrera, Flora de la Provincia de Jujuy IX, Verbénaceas a Caliceraceas. INTA, Buenos Aires.Google Scholar
    • Epling, C. & Játiva, C. (1964). Revisión del genero Satureja en América del Sur. Brittonia 16: 393 – 416.CrossRefGoogle Scholar
    • Epling, C. (1937). Synopsis of South American Labiatae. Repert. Spec. Nov. Regni Veg. 85: 97 – 192.Google Scholar
    • Epling, C. (1927). Studies in South American Labiatae III. Ann. Missori Bot. Gard. 14: 47 – 86.CrossRefGoogle Scholar
    • Fries, R. E. (1905). Zur Kenntnis der alpinen Flora in nördlichen Argentinien. Nov. Acta Regiae Soc. Sci Upsal. Ser. 4: 1 – 205.Google Scholar
    • Briquet, J. (1897). Labiatae. In: A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien IV(3): 183 – 373. Wilhelm Englemann, Leipzig.Google Scholar
    • Briquet, J. (1896). Fragmenta Monographiae Labiatarum. Bull. Herb. Boiss. 4: 762 – 878.Google Scholar
    • Rusby, H. H. (1896). An enumeration of the plants collected in Bolivia by Miguel Bang [3]. Mem. Torrey. Bot. Club 6: 1 – 130.Google Scholar
    • Kuntze, C. E. O. (1891 – 1898). Revisio Generum Plantarum 2: 515 (1891); 3: 259 (1898). A. Felix, Leipzig.Google Scholar
    • Hieronymus, G. H. E. W. (1882). Plantae diaphoricae florae argentiniae. Bol. Acad. Nac. Ci. Cordova 4: 182 – 598.Google Scholar
    • Grisebach, A. H. R. (1879). Symbolae ad Floram Argentiniam. Abh. Königl. Ges. Wiss. Göttingen 24 (1): 1 – 345.Google Scholar
    • Grisebach, A. H. R. (1874). Plantae Lorentzianae. Abh. Königl. Ges. Wiss. Göttingen 19: 49 – 279.Google Scholar
    • Leybold, F. (1873). Excursión a las Pampas Argentinas. Imprenta Nacional, Santiago.Google Scholar
    • Weddell, A. A. (1857). Chloris Andina. Vol. 2. P. Bertrand, Paris.Google Scholar
    • Bentham, G. (1832 – 5). Labiatarum Genera et Species. London, 1 – 60 (1832), 61 – 322 (1833), 323 – 645 (1834), 646 – 783 (1835).Google Scholar

    Kew Backbone Distributions

    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741.


    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 World Checklist of Selected Plant Families.

    Kew Bulletin
    Kew Bulletin

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at and
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.