1. Family: Sphenocleaceae T.Baskerv.
    1. Genus: Sphenoclea Gaertn.
      1. Sphenoclea zeylanica Gaertn.

        This species is accepted, and its native range is Africa, Tropical Asia.

    [FZ]

    Sphenocleaceae, M. Thulin. Flora Zambesiaca 7:1. 1983

    Habit
    Herb, up to 1·5 m. tall.
    Stem
    Stem hollow, often much branched.
    Leaves
    Leaf–blade lanceolate to narrowly ovate or elliptic, up to 12 x 5 cm., attenuate at the base, acute or subacute at the apex; petiole up to 1·5 cm. long.
    Inflorescences
    Spikes cylindrical–conical, up to 12 cm. long but usually much shorter, and c. 1 cm. in diameter, with only a few flowers open at a time; peduncle up to 8 cm. long; bracts and bracteoles oblanceolate–spathulate, the tips arched over the flowers except at anthesis.
    Flowers
    Flowers sessile, attached longitudinally to the rhachis by a linear base.
    Calyx
    Calyx–lobes broadly triangular, rounded at apex, 1–1·5 mm. long.
    Corolla
    Corolla white to greenish white or pink, 2–3 mm. long; lobes ovate–triangular, united about half–way.
    Filaments
    Filaments slightly dilated at base; anthers c. 0·5 mm. long.
    Ovary
    Ovary obovoid, 1–5–2 mm. long.
    Fruits
    Capsule 4–5 mm. in diameter, dehiscing below the calyx–lobes which fall with the lid, leaving the scarious base persistent on the rhachis.
    Seeds
    Seeds yellowish brown, oblong, c. 0·5 mm. long.
    [CPLC]

    Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

    Distribution
    Nativa en Colombia; Alt. 0 - 900 m.; Amazonia, Guayana y Serranía de La Macarena, Llanura del Caribe, Orinoquia, Valle del Cauca, Valle del Magdalena.
    Habit
    Hierba
    Conservation
    Preocupación Menor
    [FTEA]

    Sphenocleaceae, H. K. Airy Shaw. Flora of Tropical East Africa. 1968

    Habit
    Roots numerous, long, cord-like.
    Stem
    Stem hollow, up to 150 cm. tall, often much branched.
    Leaves
    Leaf-blade oblong to lanceolate-oblong, attenuate at both ends, acute or obtuse, 2.5–12.5 cm. long; petiole up to 3 cm. long.
    Inflorescences
    Spikes cylindric, up to 7.5 cm. long, narrowed at apex; peduncle up to 8 cm. long.
    Bracts
    Bracts and bracteoles spatulate, the tips arched over the flowers except during anthesis.
    Flowers
    Flowers densely crowded, though characteristically only 1 or 2 open at a time, rhomboid or hexagonal by compression, sessile, wedge-shaped below, attached longitudinally to the rhachis by a linear base.
    Calyx
    Calyx-segments deltoid-semicircular, obtuse, ultimately slightly accrescent and connivent.
    Corolla
    Corolla whitish, pinkish or purplish, 2.5–4 mm. long; segments ovate-triangular, obtuse or acute, united about half-way, connivent.
    Filaments
    Filaments slightly dilated at base.
    Ovary
    Ovary obovoid, 2.5 mm. long, apex broad, free, truncate.
    Fruits
    Capsule 4–5 mm. diameter, dehiscing below the calyx-segments which fall with the lid, leaving the scarious base persistent on the rhachis.
    Seeds
    Seeds yellowish-brown, 0.5 mm. long.
    Figures
    Fig. 1.
    Habitat
    In and near pools, swamps, streamsides, periodically inundated depressions, ditches, irrigation channels, and wet places generally; 0–1250 m.
    Distribution
    widespread in tropical Africa (excluding the NE. Horn), extending south to the Transvaalalso in Madagascarwidespread (but probably introduced) in tropical Asia and America P T1 T3 T5 T6 U3 Z
    [FWTA]

    Sphenocleaceae, F. N. Hepper. Flora of West Tropical Africa 2. 1963

    Habit
    An erect glabrous herb with spongy stems 1-4 ft. high
    Flowers
    Flowers greenish-white in congested spikes
    Ecology
    In rice-fields, beside rivers and open swamps, often gregarious in tidal creeks.

    Images

    Distribution

    Doubtfully present in:

    Angola

    Native to:

    Andaman Is., Bangladesh, Benin, Botswana, Burkina, Burundi, Cameroon, Central African Repu, Chad, Congo, Egypt, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, India, Ivory Coast, KwaZulu-Natal, Madagascar, Malawi, Mali, Mauritania, Mozambique, Myanmar, Namibia, Niger, Nigeria, Northern Provinces, Rwanda, Senegal, Sierra Leone, Solomon Is., Somalia, Sri Lanka, Sudan, Swaziland, Tadzhikistan, Tanzania, Togo, Uganda, Uzbekistan, Vietnam, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Alabama, Argentina Northeast, Argentina Northwest, Belize, Bolivia, Brazil North, Brazil Northeast, Brazil Southeast, Brazil West-Central, Cambodia, Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, Guatemala, Haiti, Honduras, Lesser Sunda Is., Mexico Southwest, Nicaragua, Paraguay, Puerto Rico, Trinidad-Tobago, Venezuela

    Sphenoclea zeylanica Gaertn. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Colombia 46100.000
    Harrison, S.G. [899], Guyana 18931.000
    Jones, A.P.D. [18808], Nigeria 16965.000
    Deighton, F.C., Sierra Leone 16966.000
    Meikle, R.D., Nigeria 16967.000
    Meikle, R.D. [995], Nigeria 17394.000
    Hepper, F.N. [1261], Cameroon 18122.000
    Bidgood, S. [6061], Tanzania K000468364
    Wallich, N. [Cat. no. 1516], Malaysia K001113307
    Gomez, W. [Cat. no. 1516], Myanmar K001113310
    De Silva, F. [Cat. no. 1516] K001113306
    s.coll. [Cat. no. 1516] K001113308
    s.coll. [Cat. no. 1516] K001113309
    s.coll. [Cat. no. 1516] K001113311
    s.coll. [Cat. no. 1516] K001113312
    s.coll. [Cat. no. 1516] K001113313
    s.coll. [Cat. no. 1516], India K001113314
    s.coll. [Cat. no. 1516] K001113315
    s.coll. [Cat. no. 1516] K001113316
    s.coll. [Cat. no. 1516] K001113317

    First published in Fruct. Sem. Pl. 1: 113 (1788)

    Accepted by

    • Sennikov, A.N. (ed.) (2017). Flora of Uzbekistan 2: 1-200. Navro'z Publishers, Toshkent.
    • de Moura Júnior, E.G. & al. (2015). Updated checklist of aquatic macrophytes from Northern Brazil Acta Amazonica 45: 111-132.
    • Hammel, B.E., Grayum, M.H., Herrera, C. & Zamora, N. (eds.) (2015). Manual de Plantas de Costa Rica. Volumen VIII. Dicotyledóneas (Sabiaceae-Zygophyllaceae) Monographs in Systematic Botany from the Missouri Botanical Garden 131: 1-657. Missouri Botanical Garden.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Sikarwar, R.L.S. (2014). Angiosperm diversity assessment of Chitrakootthe legendary place of Vindhyan range, India Journal of Economic and Taxonomic Botany 38: 563-619.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
    • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
    • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Singh, M. & Ali, S.J. (2008). A study on aquatic Angiosperms of district Mau (East U.P.) Journal of East African Natural History 32: 877-890.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
    • Lê, T.C. (2005). Danh l?c các loài th?c v?t Vi?t Nam 3: 1-1248. Hà N?i : Nhà xu?t b?n Nông nghi?p.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Boulos, L. (2002). Flora of Egypt 3: 1-373. Al Hadara Publishing, Cairo.
    • Stevens, W.D., Ulloa U., C., Pool, A. & Montiel, O.M. (2001). Flora de Nicaragua Monographs in Systematic Botany from the Missouri Botanical Garden 85: i-xlii, 1-2666. Missouri Botanical Garden.
    • Balick, M. J., M. H. Nee & D.E. Atha (2000). Checklist of the vascular plants of Belize Memoirs of the New York Botanical Garden 85: 1-246.
    • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
    • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181. Missouri Botanical Garden.
    • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
    • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
    • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Launert, E. (ed.) (1983). Flora Zambesiaca 7(1): 1-394. Royal Botanic Gardens, Kew.
    • Dassanayake (ed.) (1983). A Revised Handbook to the Flora of Ceylon 4: 1-532. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
    • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
    • Airy Shaw, H.K. (1968). Flora of Tropical East Africa, Sphenocleaceae: 1-3.
    • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.

    Literature

    Flora of West Tropical Africa
    • Berhaut Fl. Sén. 197
    • Airy Shaw in Fl. Males., ser. 1, 4: 27.
    • Chev. Bot. 382
    • F.T.A. 3: 481
    • Fruct. 1: 113, t. 24, fig. 5 (1788)
    Kew Backbone Distributions
    • Sennikov, A.N. (ed.) (2017). Flora of Uzbekistan 2: 1-200. Navro'z Publishers, Toshkent.
    • Forzza, R.C., Zappi, D. & Souza, V.C. (2016-continuously updated). Flora do Brasil 2020 em construção http://reflora.jbrj.gov.br/reflora/listaBrasil/ConsultaPublicaUC/ResultadoDaConsultaNovaConsulta.do.
    • de Moura Júnior, E.G. & al. (2015). Updated checklist of aquatic macrophytes from Northern Brazil Acta Amazonica 45: 111-132.
    • Hammel, B.E., Grayum, M.H., Herrera, C. & Zamora, N. (eds.) (2015). Manual de Plantas de Costa Rica. Volumen VIII. Dicotyledóneas (Sabiaceae-Zygophyllaceae) Monographs in Systematic Botany from the Missouri Botanical Garden 131: 1-657. Missouri Botanical Garden.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Sikarwar, R.L.S. (2014). Angiosperm diversity assessment of Chitrakootthe legendary place of Vindhyan range, India Journal of Economic and Taxonomic Botany 38: 563-619.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
    • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
    • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • GBIF (2008-continuously updated). Global Biodiversity Information Facility http://www.gbif.org/.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
    • Lê, T.C. (2005). Danh l?c các loài th?c v?t Vi?t Nam 3: 1-1248. Hà N?i : Nhà xu?t b?n Nông nghi?p.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Boulos, L. (2002). Flora of Egypt 3: 1-373. Al Hadara Publishing, Cairo.
    • Stevens, W.D., Ulloa U., C., Pool, A. & Montiel, O.M. (2001). Flora de Nicaragua Monographs in Systematic Botany from the Missouri Botanical Garden 85: i-xlii, 1-2666. Missouri Botanical Garden.
    • Balick, M. J., M. H. Nee & D.E. Atha (2000). Checklist of the vascular plants of Belize Memoirs of the New York Botanical Garden 85: 1-246.
    • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
    • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181. Missouri Botanical Garden.
    • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
    • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Ovchinnikov, P.N. (ed.) (1988). Flora Tadzhikskoi SSR 9: 1-568. Izd-vo Akademii nauk SSSR, Moskva.
    • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
    • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Launert, E. (ed.) (1983). Flora Zambesiaca 7(1): 1-394. Royal Botanic Gardens, Kew.
    • Dassanayake (ed.) (1983). A Revised Handbook to the Flora of Ceylon 4: 1-532. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
    • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
    • Airy Shaw, H.K. (1968). Flora of Tropical East Africa, Sphenocleaceae: 1-3.
    • Hepper, F.N. (ed.) (1963). Flora of West Tropical Africa, ed. 2, 2: 1-544.
    • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
    Flora of Tropical East Africa
    • Roessler in Prodr. Fl. Südwestafr., 138, Sphenocleaceae (1966).
    • Hepper in Flora of West Tropical Africa, ed. 2, 2: 307, fig. 272 (1963).
    • F. W. Andr., The Flowering Plants of the Sudan 3: 71, fig. 13 (1956).
    • Airy Shaw in Fl. Males., ser. 1, 4: 27, fig. 1 (1948).
    • Robyns, Flore des Spermatophytes du Parc National Albert 2: 405 (1947).
    • Engl. in Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete, Theile C: 400 (1895).
    • W. B. Hemsl. in Flora of Tropical Africa 3: 481 (1877).
    • Gaertn., De Fruct. 1: 113, t. 24/5 (1788).

    Sources

    Art and Illustrations in Digifolia
    Digital Image © Board of Trustees, RBG Kew

    Catálogo de Plantas y Líquenes de Colombia
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0