1. Family: Asteraceae Bercht. & J.Presl
    1. Genus: Brachylaena R.Br.
      1. Brachylaena huillensis O.Hoffm.

        This species is accepted, and its native range is Kenya to S. Africa.

    [FTEA]

    Compositae, H. Beentje, C. Jeffrey & D.J.N. Hind. Flora of Tropical East Africa. 2005

    Habit
    Evergreen shrub or slender erect tree 2–40 m tall, with ascending branches and rather light crown; trunk up to ± 60 cm in diameter, usually very fluted, eventually ± buttressed; bark light grey, longitudinally fissured, fibrous, peeling in long narrow shreds; young twigs densely shortly white-tomentose, glabrescent.
    Leaves
    Leaves shortly petiolate; petiole 0.3–1 cm long; blade oblanceolate, obovate or elliptic, 3–12(–15) cm long, 1–3(–5.5) cm wide, base cuneate or rarely rounded, margins entire or sometimes remotely denticulate, apex acute to obtuse and shortly acuminate, mucronate, coriaceous, strongly discolorous, green and glabrous above, silvery white or grey with short dense persistent tomentum beneath.
    Capitulum
    Capitula subsessile, numerous in clusters on short side-branches or bracteate raceme-like axillary thyrses shorter than the leaves, the female shorter and with fewer capitula than the male; involucre ± cylindrical or campanulate, the male 2–5 mm long, the female 3–6 mm long; phyllaries 5–8, densely white-tomentose, in the female glabrescent with age, glabrous or subglabrous above, the inner much longer than the outer, the innermost 3–5 elliptic, acute, 2.5–6 mm long.
    Florets
    Florets 3–9, long-exserted; corolla white; corolla tube of male florets broader than in female, 1.3–2 mm long, lobes 5, narrowly oblong, 2–3 mm long, recurved; corolla tube of female florets slender, cylindrical, shortly 2-lipped, 5-lobed, 3.5–6 mm long, lobes slender, lanceolate, ± 1 mm long.
    Fruits
    Achenes in female columnar or flattened, slightly narrowed towards the base, 2.8–4 mm long, 5–8-ribbed, densely pubescent and glandular; pappus of pale tawny minutely barbellate setae, 4–6.5 mm long.
    Figures
    Fig. 1.
    Habitat
    Dry evergreen and semi-deciduous forest and bushland on sandy soils; 1–2000 m
    Distribution
    K4 K7 T3 T8 U3
    [FZ]

    Compositae, G. V. Pope. Flora Zambesiaca 6:1. 1992

    Habit
    A shrub or small tree to 7(30) m. tall, deciduous in dry areas; bark dark-grey, finely fissured, flaking in strips; branches becoming dark-purplish, lenticellate and narrowly sulcate, at first greyish- or brown-tomentellous later glabrescent; leaves crowded at the ends of twigs.
    Leaves
    Leaves with petioles 2–14 mm. long; lamina flat or revolute, mostly 3–10(12) x 1.2–3(4) cm., larger in coppice growth, narrowly elliptic-oblanceolate to obovate or elliptic to narrowly oblong-elliptic, acute to rounded and distinctly mucronate at the apex, cuneate below, entire, slightly to strongly revolute, occasionally in obovate leaves ± strongly spinescent-denticulate, particularly towards the apex; upper surface thinly araneose, soon glabrescent and shiny dull-green with reticulate venation subprominent; lower surface greyish- or silvery-tomentellous with prominent veins.
    Capitulum
    Capitula few to many in short spikes, racemes or subglobose clusters; male synflorescences usually contracted into dense, stalked, axillary or terminal glomerules c. 1 cm. in diam., glomerules sometimes racemosely arranged on branchlets; female synflorescences composed of numerous, stalked, 2–6-capitulate clusters in leaf axils, or clusters racemosely arranged on numerous branchlets to c. 7 cm. long. Male capitula sessile or sometimes shortly stalked, 6–7-flowered; involucres to c. 3 mm. long, cyathiform; phyllaries increasing from c. 1 mm. long and ovate outside to c. 3 mm. long inside becoming oblong, tomentose; corollas creamy-white, 3.5–5.5 mm. long, narrowly funnel-shaped with lobes c. 2 mm. long and ± recurved; pappus sparse, setae 2–3 mm. long with barbs exceeding the seta axis in width. Female capitula stalked, 5–6-flowered; involucres to c. 6 mm. long, cylindric-obconic; phyllaries increasing from c. 1 mm. long and ovate outside to c. 5 mm. long inside becoming oblong, tomentose, outer phyllaries not extending onto the capitulum stalk; corollas 4–6 mm. long, filiform, lobes erect up to c. 0.5 mm. long; achenes 3–4 mm. long, subcylindric, obscurely 6–8-ribbed, pubescent; pappus several-seriate, setae 4–5 mm. long ± terete, seta barbs ± equalling the seta axis in width.
    Male
    Male capitula sessile or sometimes shortly stalked, 6–7-flowered; involucres to c. 3 mm. long, cyathiform; phyllaries increasing from c. 1 mm. long and ovate outside to c. 3 mm. long inside becoming oblong, tomentose; corollas creamy-white, 3.5–5.5 mm. long, narrowly funnel-shaped with lobes c. 2 mm. long and ± recurved; pappus sparse, setae 2–3 mm. long with barbs exceeding the seta axis in width.
    Female
    Female capitula stalked, 5–6-flowered; involucres to c. 6 mm. long, cylindric-obconic; phyllaries increasing from c. 1 mm. long and ovate outside to c. 5 mm. long inside becoming oblong, tomentose, outer phyllaries not extending onto the capitulum stalk; corollas 4–6 mm. long, filiform, lobes erect up to c. 0.5 mm. long; achenes 3–4 mm. long, subcylindric, obscurely 6–8-ribbed, pubescent; pappus several-seriate, setae 4–5 mm. long ± terete, seta barbs ± equalling the seta axis in width.

    Images

    Distribution

    Native to:

    Angola, Botswana, Kenya, KwaZulu-Natal, Mozambique, Namibia, Northern Provinces, Tanzania, Zimbabwe

    Brachylaena huillensis O.Hoffm. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1998 Gossweiler, J. [4902], Angola K001307204
    Jan 1, 1998 Gossweiler, J. [4359], Angola K001307205
    Jan 1, 1973 Gossweiler, J. [9189], Angola K001307196
    Jan 1, 1973 Gossweiler, J. [9189], Angola K001307197
    Jan 1, 1972 Battiscombe, E. [27], Kenya K000274026 holotype
    Jan 1, 1972 Battiscombe, E. [54], Kenya K000274027 lectotype
    Gossweiler, J. [4902], Angola K001307198
    Antunes [121], Angola K000274031 Unknown type material

    First published in Bot. Jahrb. Syst. 32(1): 149 (1902)

    Accepted by

    • Roskov Y. & al. (eds.) (2018). Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.

    Literature

    Flora of Tropical East Africa
    • Beentje, Kenya Trees, Shrubs and Lianas p. 555, fig. (1994).
    • G.V. Pope in Flora Zambesiaca 6: 8 (1992).
    • Paiva in Bol. Soc. Brot., sér. 2A, 46: 369 (1972).
    • O. Hoffm. in A. Engler, Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 32: 149 (1892).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0