1. Family: Fabaceae Lindl.
    1. Uraria Desv.

      1. This genus is accepted, and its native range is Tropical & Subtropical Old World.

    [LOWO]

    Legumes of the World. Edited by G. Lewis, B. Schrire, B. MacKinder & M. Lock. Royal Botanic Gardens, Kew. (2005)

    Note

    The tribe Desmodieae as treated by Ohashi et al. (1981) comprised 27 genera and c. 540 species in three subtribes, the Bryinae, Desmodiinae and Lespedezinae. Molecular analyses by Bailey et al. (1997) and Doyle et al. (2000) show that Bryinae has affinities elsewhere; Lavin et al. (2001a) place it within the Pterocarpus clade of the Dalbergieae sens. lat. (see page 309). The Bryinae are therefore removed from the Desmodieae here, as are two genera formerly placed in subtribe Lespedezinae; Phylacium Benn. and Neocollettia Hemsl., which are moved to tribe Phaseoleae (see page 393) on morphological, palynological and molecular evidence (Doyle et al., 2000; Kajita et al., 2001). The two remaining subtribes of Desmodieae are recognised in this treatment as three groups, the Lespedeza, Phyllodium and Desmodium groups, based on results of an analysis of the chloroplast gene rbcL (Kajita et al., 2001). The Phyllodium and Desmodium groups correspond to subtribe Desmodiinae, and the Lespedeza group to subtribe Lespedezinae (with Campylotropis now comprising 37 instead of 65 species as in Ohashi et al., 1981).

    Desmodieae as circumscribed here comprises 30 genera and (524)–527–(530) species (Fig. 48). The tribe occurs in the tropical, subtropical and warm temperate regions of the world, but extends into the cool temperate and sub-boreal regions of E Asia and N America (except W of the Rocky Mountains). At generic level subtribe Desmodiinae is most diverse in tropical S and SE Asia (Dy Phon et al., 1994), while temperate E Asia (Yang & Huang, 1995) and N America (Isely, 1998) are the centres of diversity of subtribe Lespedezinae. The tribe occurs widely from coastal to montane areas, but not at high altitudes. Species are most commonly shrubs or subshrubs, sometimes herbs, rarely trees and are usually erect and 3-foliolate.

    The Desmodieae have been considered similar to tribe Phaseoleae (Polhill, 1981a) and were recently shown to be a monophyletic lineage included within Phaseoleae sens. lat. (Fig. 47, page 394), closely related to subtribe Kennediinae (Doyle & Doyle, 1993, Bruneau et al., 1995; Doyle et al., 1997) and possibly sister to Mucuna (Bailey et al., 1997; Doyle et al., 2000; Kajita et al., 2001).

    Habit
    Subshrubs or shrubs
    Ecology
    Seasonally dry tropical woodland or grassland
    Distribution
    India, Indo-China, S China, Taiwan, Malesia, and two extending to Africa with the centre of diversity in India to Indo-China
    [FTEA]

    Leguminosae, J. B. Gillett, R. M. Polhill & B. Verdcourt. Flora of Tropical East Africa. 1971

    Habit
    Perennial herbs or subshrubs, prostrate or erect
    Leaves
    Leaves pinnately 3–9-foliolate, less often partly or all 1-foliolate; leaflets often large and venose; stipules persistent, free, acuminate; stipels present
    Inflorescences
    Inflorescences mostly terminal, spike-like racemes or panicles; primary bracts ovate or lanceolate, persistent or deciduous; secondary bracts and bracteoles absent
    Calyx
    Calyx 5-lobed; 3 lower lobes equal, and longer than the upper pair (often appearing to be lower pair due to twisting)
    Corolla
    Corolla yellowish or purplish; standard rounded or obovate, narrowed into a claw; wings oblong-falcate, adhering to the keel, which is slightly incurved and obtuse
    Stamens
    Vexillary stamen free; anthers uniform
    Pistil
    Ovary sessile or shortly stipitate, 2–many-ovuled; style filiform, recurved at apex; stigma terminal, capitate
    Fruits
    Pods folded like a concertina, mostly enclosed in the persistent calyx, subsessile, constricted between the seeds, the segments ovate, inflated, 1-seeded and indehiscent
    Seeds
    Seeds subglobose or compressed, oblong-ellipsoid; hilum lateral, aril not developed.
    [FZ]

    Leguminosae, B. Verdcourt. Flora Zambesiaca 3:6. 2000

    Habit
    Perennial herbs or subshrubs, prostrate or erect.
    Leaves
    Leaves pinnately 3–9-foliolate, less often partly or all 1-foliolate; leaflets often large and venose; stipules persistent, free, acuminate; stipels present.
    Inflorescences
    Inflorescences mostly terminal spike-like racemes or panicles; primary bracts ovate or lanceolate, persistent or deciduous; secondary bracts and bracteoles absent.
    Calyx
    Calyx 5-lobed; 3 lower lobes equal, and longer than the upper pair (often appearing to be lower pair due to twisting).
    Corolla
    Corolla yellowish or purplish; standard rounded or obovate, narrowed into a claw; wings oblong-falcate, adhering to the keel, which is slightly incurved and obtuse.
    Stamens
    Vexillary stamen free; anthers uniform.
    Pistil
    Ovary sessile or shortly stipitate, 2–many-ovuled; style filiform, recurved at apex; stigma terminal, capitate.
    Fruits
    Fruit folded like a concertina, mostly enclosed in the persistent calyx, subsessile, constricted between the seeds, the segments ovate, inflated, 1-seeded and indehiscent.
    Seeds
    Seeds subglobose or compressed, oblong-ellipsoid; hilum lateral, aril not developed.
    [LOWO]
    Use
    Some species used for medicine; seeds used in human food

    Images

    Distribution

    Native to:

    Andaman Is., Angola, Assam, Bangladesh, Benin, Bismarck Archipelago, Borneo, Burkina, Cambodia, Cameroon, Central African Repu, Chad, China North-Central, China South-Central, China Southeast, Congo, East Himalaya, Ethiopia, Fiji, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Hainan, India, Ivory Coast, Jawa, Laos, Lesser Sunda Is., Liberia, Malawi, Malaya, Mali, Maluku, Mozambique, Myanmar, Nansei-shoto, Nepal, New Caledonia, New Guinea, New South Wales, Nicobar Is., Niger, Nigeria, Northern Territory, Pakistan, Philippines, Queensland, Rwanda, Samoa, Senegal, Sierra Leone, Sri Lanka, Sudan, Sulawesi, Sumatera, Taiwan, Tanzania, Thailand, Tibet, Togo, Uganda, Vanuatu, Vietnam, Wallis-Futuna Is., West Himalaya, Western Australia, Zaïre

    Introduced into:

    Niue, Tonga

    Uraria Desv. appears in other Kew resources:

    First published in J. Bot. Agric. 1: 122 (1813)

    Literature

    Flora of West Tropical Africa
    • —F.T.A. 2: 168.
    Flora Zambesiaca
    • Verdcourt in Kirkia 9: 532 (1974).
    • in J. Bot. Agric. 1: 122, t. 5, fig. 19 (1813).
    Flora of Tropical East Africa
    • Journ. de Bot. 1: 122, t. 5/19 (1813)

    Sources

    Art and Illustrations in Digifolia
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Legumes of the World Online
    http://creativecommons.org/licenses/by-nc-sa/3.0