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This species is accepted, and its native range is Bolivia to Brazil and N. Argentina.
A specimen from Kew's Herbarium

[FZ]

Flora Zambesiaca. Vol 8, Pt 7. Avicenniaceae, R. Fernandes. Nesogenaceae, M.A. Diniz. Verbenaceae, R. Fernandes. Lamiaceae, R. Fernandes. 2005.

Type
Type from Paraguay. Type from Brazil.
Morphology General
Perennial prostrate trailing herb 10–80 cm high
Morphology Reproductive morphology Flowers Calyx
Calyx 7.5–9 mm long, tubular, becoming wider below and constricted above in fruit, densely appressed whitish strigose-pubescent with some intermixed darker glands; teeth ovate at the base, and abruptly contracted into a long seta 0.75–2 mm long, unequal Calyx 7.5–9 mm long, tubular, becoming wider below and constricted above in fruit, densely appressed whitish strigose-pubescent with some intermixed darker glands; teeth ovate at the base, and abruptly contracted into a long seta 0.75–2 mm long, unequal.
Morphology Reproductive morphology Flowers Corolla
Corolla mauve, lilac, mauve-blue, purple, purple-red, deep pink or sometimes white; tube 10–11 mm long, exserted for 3–5 mm from the calyx mouth, glabrous outside, hairy inside at the throat, with two longitudinal dense bands of white setae below the insertions of the lower stamens; limb c. 10 mm wide, the lobes 3–4 × 3–4 mm, broadly obcordate, emarginate Corolla mauve, lilac, mauve-blue, purple, purple-red, deep pink or sometimes white; tube 10–11 mm long, exserted for 3–5 mm from the calyx mouth, glabrous outside, hairy inside at the throat, with two longitudinal dense bands of white setae below the insertions of the lower stamens; limb c. 10 mm wide, the lobes 3–4 × 3–4 mm, broadly obcordate, emarginate.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens included, or with glands just exserted Stamens included, or with glands just exserted.
Morphology Reproductive morphology Flowers Gynoecium Style
Style 9–13 mm long, included, persistent Style 9–13 mm long, included, persistent.
Morphology Reproductive morphology Fruits
Mericarps c. 3.5 mm long, black, striate, longitudinally raised-reticulate in upper part. Mericarps c. 3.5 mm long, black, striate, longitudinally raised-reticulate in upper part.
Ecology
In Brachystegia woodland and grassland; also in old cultivation and on roadsides; 100–1520 m.
Note
Although Moldenke, in Phytologia 8 : 190 (1962) and Phytologia 11 : 284 (1965), treated V. tenuisecta Briq. and V. aristigera S. Moore as separate species, it is here considered best to follow Troncoso who, in Darwiniana 14 : 636 (1968), considered them to be synonymous.
Distribution
Botswana Zimbabwe Malawi Mozambique Native of South America (Brazil, Uruguay, Argentina, Paraguay, etc.), now widely cultivated as a garden ornamental, often becoming naturalized.  In Africa it also occurs in Egypt and Nigeria (fide Moldenke ), in Dem. Rep. Congo, South Africa (KwaZu District code: BOT SE, BOT W, ZIM W, ZIM C, ZIM E, MAL S, MOZ GI, MOZ M.
Morphology General Habit
Perennial prostrate trailing herb 10–80 cm high.
Morphology Stem
Stems often woody at the base, decumbent, sometimes rooting at the lower nodes, tetragonal, sparsely pilose to pubescent or glabrescent. Stems often woody at the base, decumbent,  sometimes rooting at the lower nodes, tetragonal, sparsely pilose to pubescent or glabrescent
Morphology Leaves
Leaves petiolate, with short sterile densely leafy branchlets or fascicles of smaller leaves in the axils, 2–3.5 × 2–3 cm, ± triangular or ovate in outline, 2-pinnatisect or 3-sect with the segments also pinnatisect or 3-sect; the ultimate segments up to 1 cm long and c. 1 mm wide, linear, obtuse to acute; lamina somewhat thick, revolute on the margins, strigose-pubescent or ± glabrous on both surfaces, sometimes with brownish glands, entire or toothed; petiole up to 2 cm long. Leaves petiolate, with short sterile densely leafy branchlets or fascicles of smaller leaves in the axils, 2–3.5 × 2–3 cm, ± triangular or ovate in outline, 2-pinnatisect or 3-sect with the segments also pinnatisect or 3-sect; the ultimate segments up to 1 cm long and c. 1 mm wide, linear, obtuse to acute; lamina somewhat thick, revolute on the margins, strigose-pubescent or ± glabrous on both surfaces, sometimes with brownish glands, entire or toothed; petiole up to 2 cm long
Morphology Reproductive morphology Inflorescences
Inflorescences of solitary terminal spikes on stems and branches; spikes c. 1.5 × 2 cm, ovoid and dense in flower, elongating to 5–12 cm in fruit; peduncles 1.5–5 cm long, elongating to 10 cm or more in fruit; bracts green, 1/3 to 1/2 as long as the calyx, ovate-lanceolate, densely canescent-puberulous or strigose and sometimes glandular; flowers numerous showy, the upper ones dense and ascending, the lower ones spreading. Inflorescences of solitary terminal spikes on stems and branches; spikes c. 1.5 × 2 cm, ovoid and dense in flower, elongating to 5–12 cm in fruit; peduncles 1.5–5 cm long, elongating to 10 cm or more in fruit; bracts green, 1/3 to1/2 as long as the calyx, ovate-lanceolate, densely canescent-puberulous or strigose and sometimes glandular; flowers numerous showy, the upper ones dense and ascending, the lower ones spreading

[FTEA]

Verbenaceae, B. Verdcourt. Flora of Tropical East Africa. 1992

Morphology General Habit
Perennial herb, often woody at the base, 10–40 cm. tall; stems decumbent below, sparsely pubescent but soon glabrescent.
Morphology Leaves
Leaves ± triangular in oudine, pinnatisect, 2–3.5 cm. long, 2–4 cm. wide; segments linear, up to 1 cm. long, 0.5–1(–3) mm. wide, ± acute or obtuse at the apex, substrigose-pubescent or ± glabrous, entire or toothed.
Morphology Reproductive morphology Inflorescences
Inflorescences terminal, dense and ± ovate, at first 1.5 × 2 cm. but soon elongating to 5–12 cm.; peduncles 1.5–5 cm. long, elongating to 10 cm. or more in fruit.
Morphology Reproductive morphology Inflorescences Bracts
Bracts subovate at base, lanceolate-subulate, 2–4 mm. long, 1.2–1.5 mm. wide, canescent-puberulous.
Morphology Reproductive morphology Flowers Calyx
Calyx tubular, 6–7 mm. long, densely adpressed strigose-pubescent and with some sparse dark glands; teeth ovate at base, contracted into along seta, 0.5–2 mm. long, unequal.
Morphology Reproductive morphology Flowers Corolla
Corolla blue or purple; tube 1–1.1 cm. long, glabrous outside, hairy inside at throat; limb ± 1 cm. wide, the lobes ± broadly obcordate, 3–4 mm. long and wide, emarginate for 1–1.5 mm.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens included or with glands just exserted.
Morphology Reproductive morphology Flowers Gynoecium Style
Style ± included, 0.9–1.3 cm. long, persistent.
Morphology Reproductive morphology Fruits
Nutlets subcylindrical, 3 mm. long, 0.8 mm. wide, reticulately ribbed outside, white verrucose inside.
Habitat
Cultivated but also as a weed locally on cleared land; 1650–2060 m.
Distribution
K3 K4 T2

Native to:

Argentina Northeast, Argentina Northwest, Bolivia, Brazil South, Brazil Southeast, Brazil West-Central, Paraguay, Uruguay

Introduced into:

Botswana, California, Free State, Greece, Guatemala, Honduras, India, Kenya, KwaZulu-Natal, Lesotho, Malawi, Mozambique, New South Wales, Nicaragua, Nigeria, Northern Provinces, Northern Territory, Queensland, South Australia, Swaziland, Tanzania, Venezuela, Victoria, Western Australia, Zambia, Zaïre, Zimbabwe

Verbena aristigera S.Moore appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Dec 1, 2008 Huaylla, A. [2065], Bolivia Glandularia aristigera K000509775
Balansa, B. [1025], Paraguay Verbena tenuisecta K000470520 Unknown type material

First published in Trans. Linn. Soc. London, Bot. 4: 439 (1895)

Accepted by

  • Retief, E. & Meyer, N.L. (2017). Plants of the Free State inventory and identification guide Strelitzia 38: 1-1236. National Botanical Institute, Pretoria.

Not accepted by

  • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew. [Cited as Glandularia aristigera.]

Literature

Flora Zambesiaca

  • Cabrera, Man. Fl. Alreded. Buenos Aires: 397 (1953).
  • Darwiniana 13 : 477, fig. 4 (1964).
  • F.T.E.A., Verbenaceae: 10 (1992).
  • J.H. Ross, Fl. Natal: 299 (1972).
  • Phytologia 11 , 4: 280 (1965)
  • Phytologia 8 : 189 (1952).
  • Rev. Fl. Ceylon 4 : 208 (1983).
  • op. cit. 16 : 208 (1968)
  • op. cit. 41 : 403 (1979).

Kew Backbone Distributions

  • Davidse, G. & al. (eds.) (2012). Flora Mesoamericana 4(2): 1-533. Universidad Nacional Autónoma de México, México, D.F.
  • Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A., & Tzanoudakis, D. (2013). Vascular plants of Greece. An annotated checklist: 1-372. Botanic gardens and botanical museum Berlin-Dahlem, Berlin and Hellenic botanical society, Athens.
  • Fernandes, R. & Diniz, M.A. (2005). Avicenniaceae, Nesogenaceae, Verbenaceae and Lamiaceae (subfams, Viticoideae and Ajugoideae) Flora Zambesiaca 8(7): 1-161. Royal Botanic Gardens, Kew.
  • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
  • O'Leary, N. & Thode, V. (2016). The genus Glandularia (Verbenaceae) in Brazil Annals of the Missouri Botanical Garden 101: 699-749.
  • Peralta, P.F. & Múlgura, M.E. (2011). El género Glandularia (Verbenaceae) en Argentina Annals of the Missouri Botanical Garden 98: 358-412.
  • Retief, E. & Meyer, N.L. (2017). Plants of the Free State inventory and identification guide Strelitzia 38: 1-1236. National Botanical Institute, Pretoria.

Flora of Tropical East Africa

  • Moldenke in Phytologia 8: 189 (1962).
  • S. Moore in Trans. Linn. Soc., Bot., ser. 2, 4: 439 (1895).

Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Neotropikey
Milliken, W., Klitgard, B. and Baracat, A. (2009 onwards), Neotropikey - Interactive key and information resources for flowering plants of the Neotropics.
http://creativecommons.org/licenses/by/3.0