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This species is accepted, and its native range is Tropical & Subtropical Old World, NE Queensland.

[FZ]

Cucurbitaceae, C. Jeffrey. Flora Zambesiaca 4. 1978

Habit
Stems prostrate or scandent to 5 m., sparsely to densely crispate-pubescent or villous, especially at nodes.
Leaves
Leaf-lamina 1–10 × 1–12·5 cm., broadly ovate to orbicular in outline, cordate, narrowly decurrent on to petiole, punctate and sparsely pubescent to densely villous on veins beneath, sparsely hirsute especially on veins above, deeply palmately 3–7-lobed, lobes variously sinuate-dentate or lobulate, acute to retuse, apiculate.
Petiole
Petiole 0·5–7 cm. long.
Tendrils
Tendrils simple.
Flowers
Flowers monoecious, solitary. male flowers: peduncle 0·3–5 cm. long; bract 2–17 mm. long, broadly ovate or reniform, sessile, cordate, amplexicaul, obtuse or retuse, apiculate, green; pedicel 2–9·5 cm. long. Receptacle-tube 1–5 mm. long; lobes 3–7 mm. long, ovate-lanceolate. Petals 1·0–2·5 cm. long, pale to deep yellow, ovate to obovate. Female flowers: peduncle 0·2–5 cm. long; bract 1–12 mm. long; pedicel 1–10 cm. long; ovary 8–11(30) × 2–4 mm., ovoid-rostrate to fusi-form, ridged, pilose on ridges, tuberculate; receptacle-tube 1–3 mm. long, lobes 2–5 mm. long, lanceolate; petals 0·7–1·2 cm. long.
Male
male flowers: peduncle 0·3–5 cm. long; bract 2–17 mm. long, broadly ovate or reniform, sessile, cordate, amplexicaul, obtuse or retuse, apiculate, green; pedicel 2–9·5 cm. long. Receptacle-tube 1–5 mm. long; lobes 3–7 mm. long, ovate-lanceolate. Petals 1·0–2·5 cm. long, pale to deep yellow, ovate to obovate.
Female
Female flowers: peduncle 0·2–5 cm. long; bract 1–12 mm. long; pedicel 1–10 cm. long; ovary 8–11(30) × 2–4 mm., ovoid-rostrate to fusi-form, ridged, pilose on ridges, tuberculate; receptacle-tube 1–3 mm. long, lobes 2–5 mm. long, lanceolate; petals 0·7–1·2 cm. long.
Fruits
Fruit 2·5–4·8(11) × 1·5–2·3(4) cm., ovoid-rostrate or ellipsoid, longitudinally ribbed and tuberculate, bright orange-red, dehiscent into 3 valves; fruit-stalk 3·4–15 cm. long.
Seeds
Seeds 8–11 × 4·5–8 × 2–3·5 mm., enveloped in sticky red pulp, ovate-elliptic to oblong in outline; faces flattened, sculptured, with sinuate edges; margins grooved.

[FTEA]

Cucurbitaceae, C. Jeffrey. Flora of Tropical East Africa. 1967

Habit
Annual climber or trailer to 5 m.
Stem
Stems herbaceous, ridged, glabrous or hairy, strap-shaped at the base.
Leaves
Leaf-blade broadly ovate-reniform or orbicular in outline, cordate, 25–100 mm. long, 30–125 mm. broad, glabrous or pubescent (especially on the nerves) on both surfaces, deeply palmately (3–)5(–7)-lobed; lobes obovate, ovate-elliptic or rhombic in outline, narrowed towards the base, acute and apiculate, sinuate-toothed and unlobed or more usually sinuate-lobulate; petioles 14–70 mm. long.
Tendrils
Tendrils simple.
Flowers
Monoecious. Male flowers solitary; peduncle 5–28 mm. long, bearing an apical 3–15 mm. long 4–22 mm. broad suborbicular apiculate clasping green bract; pedicel 18–55 mm. long; receptacle-tube obconic, pale greenish-yellow, 2–4 mm. long; lobes ovate-lanceolate or elliptic, adpressed to lower surface of the petals, 3–6.5 mm. long; petals obovate-lingulate, apiculate, 11–22 mm. long, 7.5–15 mm. broad, pale yellow to orange-yellow, 2 with scales inside at the base; stamens 3; anthers coherent in centre of flower; thecae triplicate. Female flowers with peduncle 2–50 mm. long, bearing an apical broadly ovate or suborbicular 1–12 mm. long 1–10 mm. broad sessile bract; pedicel 10–100 mm. long; ovary ovoid and beaked to elongate-fusiform, muricate-tuberculate, 8–30 mm. long, 2–4 mm. across; receptacle-lobes linear, acute, 2–5 mm. long, reflexed at the tips; petals 7–12 mm. long, 3–6 mm. wide.
Male
Male flowers solitary; peduncle 5–28 mm. long, bearing an apical 3–15 mm. long 4–22 mm. broad suborbicular apiculate clasping green bract; pedicel 18–55 mm. long; receptacle-tube obconic, pale greenish-yellow, 2–4 mm. long; lobes ovate-lanceolate or elliptic, adpressed to lower surface of the petals, 3–6.5 mm. long; petals obovate-lingulate, apiculate, 11–22 mm. long, 7.5–15 mm. broad, pale yellow to orange-yellow, 2 with scales inside at the base; stamens 3; anthers coherent in centre of flower; thecae triplicate.
Female
Female flowers with peduncle 2–50 mm. long, bearing an apical broadly ovate or suborbicular 1–12 mm. long 1–10 mm. broad sessile bract; pedicel 10–100 mm. long; ovary ovoid and beaked to elongate-fusiform, muricate-tuberculate, 8–30 mm. long, 2–4 mm. across; receptacle-lobes linear, acute, 2–5 mm. long, reflexed at the tips; petals 7–12 mm. long, 3–6 mm. wide.
Fruits
Fruit on a 37–150 mm. or more long stalk, pendulous, broadly ovoid and beaked to attenuate-ellipsoid, 35–110 mm. long, 20–40 mm. across, reddish-orange with a paler apex, ornamented with ± 8 longitudinal rows of subconical tubercles and with many smaller tubercles in between, splitting into 3 valves and exposing the seeds sheathed in sticky red pulp hanging in two rows from the faces of each valve.
Seeds
Seeds oblong, ± 8–11 × 4.5–6 × 2.5–3.5 mm.; testa sculptured; margins grooved.
Habitat
Lowland rain-forest and riverine forest, elephant-grass thicket and in plantations; 100–1220 m.; sometimes cultivated
Distribution
K7 T4 T8 U2 U4 Z pantropical, but probably introduced in the New World

[CPLC]

Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

Distribution
Adventicia en Colombia; Alt. 5 - 1500 m.; Amazonia, Andes, Islas Caribeñas, Llanura del Caribe, Orinoquia, Pacífico, Valle del Cauca, Valle del Magdalena.
Habit
Trepadora
Conservation
No Evaluada

[FWTA]

Cucurbitaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:1. 1954

Habit
Climber, flowers yellow
Fruits
Fruit ripening orange, 2–3 in. long, bursting in 3 valves to expose the carmine red seeds
Ecology
Extending into moister regions.

Native to:

Angola, Assam, Bangladesh, Benin, Borneo, Burkina, Burundi, Cameroon, Cape Verde, Central African Repu, Chad, Christmas I., Congo, East Himalaya, Ethiopia, Fiji, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., India, Ivory Coast, Jawa, Kenya, KwaZulu-Natal, Laos, Lesser Sunda Is., Liberia, Malawi, Malaya, Mali, Maluku, Mozambique, Namibia, New Guinea, Niger, Nigeria, Northern Provinces, Northern Territory, Philippines, Queensland, Rwanda, Senegal, Sierra Leone, Society Is., Solomon Is., Sri Lanka, Sudan, Sulawesi, Sumatera, Tanzania, Thailand, Tonga, Uganda, Vietnam, West Himalaya, Western Australia, Zambia, Zaïre, Zimbabwe

Extinct in:

Socotra

Introduced into:

Alabama, Andaman Is., Argentina Northeast, Argentina Northwest, Aruba, Ascension, Bahamas, Belize, Bermuda, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Cambodia, Cayman Is., Central American Pac, China South-Central, China Southeast, Colombia, Connecticut, Cook Is., Costa Rica, Cuba, Dominican Republic, El Salvador, Florida, Galápagos, Georgia, Guatemala, Haiti, Honduras, Jamaica, Korea, Leeward Is., Louisiana, Marianas, Mauritius, Mexico Northwest, Mexico Southwest, Netherlands Antilles, New Caledonia, Nicobar Is., Niue, Panamá, Paraguay, Pennsylvania, Puerto Rico, Rodrigues, Réunion, Samoa, Southwest Caribbean, Trinidad-Tobago, Turks-Caicos Is., Venezuela, Venezuelan Antilles, Windward Is.

Momordica charantia L. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Jan 1, 2016 Hooker, J.D. [162], Cape Verde K001134350
Jan 1, 2009 Fernandes, D. [464], Bolivia K000634046
Oct 3, 2007 Hamilton, M.A. [185], Montserrat 76505.000
Jan 1, 2006 Cheek, M. [11710], Cameroon K000582255
Feb 1, 2003 Cheek, M. [5634], Cameroon K000008307
Feb 1, 2003 Cheek, M. [7524], Cameroon K000008305
Feb 1, 2003 Cheek, M. [7524], Cameroon K000008306
Feb 1, 2003 Cable, S. [3231], Cameroon K000008308
Feb 1, 2003 Cable, S. [2799], Cameroon K000008301
Feb 1, 2003 Sidwell, K. [385], Cameroon K000008303
Feb 1, 2003 Zapfack, L. [642], Cameroon K000008302
Dec 1, 1967 Leeuwenberg, A.J.M. [6945], Cameroon K000050716
22677.000
29047.008
4646.000
Milne-Redhead, E. [7596], Tanzania 5158.000
Verdcourt, B. [3942], Kenya 23968.000
Wild, H. [44470], Zimbabwe 823.000
Tweedie, E.M. [3484], Kenya 32458.000
Bullock, A.A. [3999], Zambia 6519.000
Purseglove, J.W. [3258], Uganda 8104.000
Richards, H.M. [16056], Tanzania 4994.000
Harley, R.M. [16272], Brazil 35923.000
Harley, R.M. [16272], Brazil K001139632
Harley, R.M. [16212], Brazil K001139633
Harley, R.M. [17255], Brazil K001139634
White, F. [2379], Zambia 8105.000
Hunt, D.R. [6162], Brazil K001139667
Philcox, D. [4039], Brazil K001139652
Philcox, D. [4039], Brazil K001139653
Philcox, D. [3860], Brazil K001139654
Trail, J.W.H. [362], Brazil K001139617
Hutchinson, J. [99], Cameroon K000050718
Hutchinson, J. [99], Cameroon K000050719
Hutchinson, J. [99], Cameroon K000050720
Harley, R. [5182], Brazil K001139624
Harley, R. [5182], Brazil K001139625
Radcliffe-Smith, A. [3882], Oman 38880.000
Hatschbach, G. [17173], Brazil K001139666
Hatschbach, G. [45119], Brazil K001139609
Hatschbach, G. [38552], Brazil K001139619
Hatschbach, G. [33807], Brazil K001139620
Hatschbach, G. [5203], Brazil K001139621
Hatschbach, G. [49168], Brazil K001139627
Hatschbach, G. [31372], Brazil K001139629
Hatschbach, G. [29505], Brazil K001139630
Rico, L. [1415], Bolivia K000295406
Lindeman, J.C. [4573], Brazil K001139678
Pearson, H.P.N. [4], Brazil K001139649
Silva, J.M. [1930], Brazil K001139676
Hage, J.L. [589], Brazil K001139614
Irwin, H.S. [21484], Brazil K001139665
Irwin, H.S. [15866], Brazil K001139668
Andrade, I.M. [84], Brazil K001139607
Agra, M.F. [1169], Brazil K001139679
Eiten, G. [4571], Brazil K001139651
Preston, T.A. [s.n.], Brazil K001139661
Preston, T.A. [s.n.], Brazil K001139662
Preston, T.A. [s.n.], Brazil K001139663
Lemos, M.J.S. [20], Brazil K001139677
Lemos, M.J.S. [20], Brazil K001139622
França, F. [2290], Brazil K001139623
Pickersgill, B. [Ru71-272], Brazil K001139616
Maas, P.J.M. [322A], Brazil K001139674
Bolland, G. [s.n.], Brazil K001139640
Kummrow, R. [538], Brazil K001139615
Kummrow, R. [508], Brazil K001139618
Silva, N.T. [2059], Brazil K001139673
Riedel [707], Brazil K001139648
Burchell, W.J. [7158-2], Brazil K001139660
Miers, J. [3491], Brazil K001139647
Robert, A. [748], Brazil K001139655
Graham [s.n.], Brazil K001139639
Moseley, H.N. [s.n.], Cape Verde K001134351
Teixeira, L.O.A. [378], Brazil K001139610
Teixeira, L.O.A. [720], Brazil K001139611
Daly, D.C. [D520], Brazil K001139612
Balée, W.L. [1542], Brazil K001139608
Vieira, G. [389], Brazil K001139635
Prance, G.T. [4004], Brazil K001139669
Prance, G.T. [4004], Brazil K001139670
Prance, G.T. [14673], Brazil K001139671
Prance, G.T. [4004], Brazil K001139631
Lowrie, S.R. [400], Brazil K001139613
Balick, M.J. [1545], Brazil K001139650
Traill, J.W.H. [362], Brazil K001139638
Mexia, Y. [4543], Brazil K001139664
s.coll. [s.n.] K000742645
s.coll. [s.n.], India K000742644
Manning, S.D. [1265], Cameroon K000050717
Lobo, M.G.A. [156], Brazil K001139636
Swainson, W. [s.n.], Brazil K001139644
Albuquerque, B.W. [1363], Brazil K001139672
Vieria, R.F. [93], Brazil K001139628
Lobão, A.Q. [28], Brazil K001139641
Gibert, E.J. [37], Paraguay K001139645
s.coll. [Cat. no. 6746] K001124719
s.coll. [s.n.] K001139643
s.coll. [s.n.], Brazil K001139646
Burchell [822], Brazil K001139658
Burchell [7158-2], Brazil K001139659
Fonseca, V.S. [116], Brazil K001139642
Moseley [s.n.], Brazil K001139656
Moseley [s.n.], Brazil K001139657
Damineli, E.C. [20426], Brazil K001139675
Skorupa, L. [90], Brazil K001139626
Döbereiner, J. [1470], Brazil K001139637
Milne-Redhead, E. [5140], Ghana Momordica charantia var. abbreviata 4645.000
s.coll. [Cat. no. 6745] Momordica muricata K001124713
s.coll. [Cat. no. 6745] Momordica muricata K001124714
s.coll. [Cat. no. 6745] Momordica muricata K001124715
s.coll. [Cat. no. 6745] Momordica muricata K001124716
s.coll. [Cat. no. 6745] Momordica muricata K001124717
s.coll. [Cat. no. 6745] Momordica muricata K001124718

First published in Sp. Pl.: 1009 (1753)

Accepted by

  • Zuloaga, F.O. & Belgrano, M.J. (eds.) (2017). Flora Argentina. Flora vascular de la República Argentina 17: 1-434. INTA, IMBIV & IBODA.
  • de Moura Júnior, E.G. & al. (2015). Updated checklist of aquatic macrophytes from Northern Brazil Acta Amazonica 45: 111-132.
  • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
  • Flora of North America Editorial Committee (2015). Flora of North America North of Mexico 6: 1-468. Oxford University Press, New York, Oxford.
  • Sikarwar, R.L.S. (2014). Angiosperm diversity assessment of Chitrakootthe legendary place of Vindhyan range, India Journal of Economic and Taxonomic Botany 38: 563-619.
  • Velayos, M., Barberá, P., Cabezas, F.J., de la Estrella, M., Fero, M. & Aedo, C. (2014). Checklist of the vascular plants of Annobón (Equatorial Guinea) Phytotaxa 171: 1-78.
  • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
  • Garcillán, P.P. & al. (2013). Plantas no nativas naturalizadas de la península de Baja California, México Botanical Sciences 91: 461-475.
  • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
  • Rebnner, S.S. & Pandey, A.K. (2013). The Cucurbitaceae of India: Accepted names, synonyms, geographic distribution, and information on images and DNA sequences PhytoKeys 20: 53-118.
  • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
  • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
  • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
  • Meena, S.L. (2012). A checklist of the vascular plants of Banaskantha district, Gujarat, India Nelumbo 54: 39-91.
  • Kumar, S. (2012). Herbaceous flora of Jaunsar-Bawar (Uttarkhand), India: enumerations Phytotaxonomy 12: 33-56.
  • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
  • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
  • Wu, Z. & Raven, P.H. (eds.) (2011). Flora of China 19: 1-884. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
  • Reddy, G.V.P. (2011). Survey of invasive plants on Guam and identification of the 20 most widespread Micronesica; Journal of the College of Guam 41: 263-274.
  • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
  • Maliya, S.D. & Datt, B. (2010). A contribution to the flora of Katarniyaghat wildlife sanctuary, Baharaich district, Uttar Pradesh Journal of Economic and Taxonomic Botany 34: 42-68.
  • Hammel, B.E., Grayum, M.H., Herrera & C. & Zamora, N. (eds.) (2010). Manual de plantas de Costa Rica volumen V. Dicotiledóneas (Clusiaceae-Gunneraceae) Monographs in Systematic Botany from the Missouri Botanical Garden 119: 1-970. Missouri Botanical Garden.
  • Araújo, D. & Alves, M. (2010). Climbing plants of a fragmented area of lowland Atlantic forest, Igarassu, Pernambuco (northeastern Brazil) Phytotaxa 8: 1-24.
  • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
  • Sarmah, K.K. & Borthakur, S.K. (2009). A checklist of angiospermic plants of Manas national park in Assam, India Pleione 3: 190-200.
  • Monteiro de Barros, A.A., de Andrade Ribas, L. & Dunn Araujo, D.S. (2009). Trepadeiras do Parque Estadual da Serra do Tiririca, Rio de Janeiro, Brasil Rodriguésia 60: 681-694.
  • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
  • de Wilde, W.J.J.O. & Duyfjes, B.E.E. (2008). Flora of Thailand 9(4): 411-546. The Forest Herbarium, Royal Forest Department.
  • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
  • De Wilde, W.J.J.O. & Duyfjes, B.E.E. (2008). The edible Cucurbitaceae of Thailand and Malesia and the wild forms of the cultivated ones Sandakania 17: 43-91.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
  • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
  • Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.
  • Trusty, J.L., Kesler, H.C. & Delgado, G.H. (2006). Vascular flora of Isla del Coco, Costa Rica Proceedings of the California Academy of Sciences, ser. 4, 57: 247-355.
  • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
  • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
  • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
  • Miller, A.G. & Morris, M. (2004). Ethnoflora of Soqotra Archipelago: 1-759. The Royal Botanic Garden, Edinburgh.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Lê, T.C. (2003). Danh l?c các loài th?c v?t Vi?t Nam 2: 1-1203. Hà N?i : Nhà xu?t b?n Nông nghi?p.
  • Grierson, A.J.C. & Long, D.G. (2001). Flora of Bhutan 2: 1-1675. Royal Botanic Gardens, Edinburgh.
  • Stevens, W.D., Ulloa U., C., Pool, A. & Montiel, O.M. (2001). Flora de Nicaragua Monographs in Systematic Botany from the Missouri Botanical Garden 85: i-xlii, 1-2666. Missouri Botanical Garden.
  • Balick, M. J., M. H. Nee & D.E. Atha (2000). Checklist of the vascular plants of Belize Memoirs of the New York Botanical Garden 85: 1-246.
  • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
  • Dassanayake (ed.) (1997). A Revised Handbook to the Flora of Ceylon 11: 1-420. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
  • Edwards, S., Tadesse, M. & Hedberg, I. (eds.) (1995). Flora of Ethiopia and Eritrea 2(2): 1-456. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Turner, I.M. (1995). A catalogue of the Vascular Plants of Malaya Gardens' Bulletin Singapore 47(1): 1-346.
  • Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
  • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
  • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
  • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
  • Zizka, G. (1991). Flowering plants of Easter island Palmarum Hortus Francofurtensis 3: 1-108.
  • Hancock, I.R. & Henderson, C.P. (1988). Flora of the Solomon Islands Research Bulletin Dodo Creek Research Station 7: 1-203.
  • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
  • Troupin, G. (ed.) (1983). Flora du Rwanda 2: 1-603. Musee Royal de l'Afrique Centrale.
  • (1982). Flora of Australia 8: 1-420. Australian Government Publishing Service, Canberra.
  • Launert, E. (ed.) (1978). Flora Zambesiaca 4: 1-658. Royal Botanic Gardens, Kew.
  • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
  • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Fosberg, F.R. (1975). Vascular plants of Aitutaki Atoll Research Bulletin 190: 73-84.
  • Keraudren-Aymonin, M. (1975). Flore d'Afrique Centrale (Zaire-rwanda-Burundi), Cucurbitaceae: 1-152. Jardin Botanique National de Belgique.
  • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Sykes, W.R. (1970). Contributions to the flora of Niue Bulletin, New Zealand Department of Scientific and Industrial Research 200: 1-321.
  • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
  • Jeffrey, C. (1967). Flora of Tropical East Africa, Cucurbitaceae: 1-156.
  • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.

Literature

Flora of West Tropical Africa

  • Cogn. & Harms in Engl. Pflanzenr. Cucurbit. 2: 24
  • Chev. Bot. 290.
  • —F.T.A. 2: 537

Kew Backbone Distributions

  • Zuloaga, F.O. & Belgrano, M.J. (eds.) (2017). Flora Argentina. Flora vascular de la República Argentina 17: 1-434. INTA, IMBIV & IBODA.
  • Jaramillo Díaz, P. & Guézou, A. (2017). CDF Checklist of Galapagos Vascular Plants - FCD Lista de especies de Plantas Vasculares de Galápagos http://www.darwinfoundation.org/datazone/checklists/vascular-plants/.
  • Forzza, R.C., Zappi, D. & Souza, V.C. (2016-continuously updated). Flora do Brasil 2020 em construção http://reflora.jbrj.gov.br/reflora/listaBrasil/ConsultaPublicaUC/ResultadoDaConsultaNovaConsulta.do.
  • de Moura Júnior, E.G. & al. (2015). Updated checklist of aquatic macrophytes from Northern Brazil Acta Amazonica 45: 111-132.
  • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
  • Flora of North America Editorial Committee (2015). Flora of North America North of Mexico 6: 1-468. Oxford University Press, New York, Oxford.
  • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
  • Garcillán, P.P. & al. (2013). Plantas no nativas naturalizadas de la península de Baja California, México Botanical Sciences 91: 461-475.
  • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
  • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
  • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
  • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
  • Kumar, S. (2012). Herbaceous flora of Jaunsar-Bawar (Uttarkhand), India: enumerations Phytotaxonomy 12: 33-56.
  • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
  • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
  • Wu, Z. & Raven, P.H. (eds.) (2011). Flora of China 19: 1-884. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
  • Reddy, G.V.P. (2011). Survey of invasive plants on Guam and identification of the 20 most widespread Micronesica; Journal of the College of Guam 41: 263-274.
  • Idárraga-Piedrahita, A., Ortiz, R.D.C., Callejas Posada, R. & Merello, M. (eds.) (2011). Flora de Antioquia: Catálogo de las Plantas Vasculares 2: 1-939. Universidad de Antioquia, Medellín.
  • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
  • de Wilde, W.J.J.O. & Duyfjes, B.B.E. (2010). Flora Malesiana 19: 1-342. Noordhoff-Kolff N.V., Djakarta.
  • Maliya, S.D. & Datt, B. (2010). A contribution to the flora of Katarniyaghat wildlife sanctuary, Baharaich district, Uttar Pradesh Journal of Economic and Taxonomic Botany 34: 42-68.
  • Hammel, B.E., Grayum, M.H., Herrera & C. & Zamora, N. (eds.) (2010). Manual de plantas de Costa Rica volumen V. Dicotiledóneas (Clusiaceae-Gunneraceae) Monographs in Systematic Botany from the Missouri Botanical Garden 119: 1-970. Missouri Botanical Garden.
  • Araújo, D. & Alves, M. (2010). Climbing plants of a fragmented area of lowland Atlantic forest, Igarassu, Pernambuco (northeastern Brazil) Phytotaxa 8: 1-24.
  • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
  • Sarmah, K.K. & Borthakur, S.K. (2009). A checklist of angiospermic plants of Manas national park in Assam, India Pleione 3: 190-200.
  • Monteiro de Barros, A.A., de Andrade Ribas, L. & Dunn Araujo, D.S. (2009). Trepadeiras do Parque Estadual da Serra do Tiririca, Rio de Janeiro, Brasil Rodriguésia 60: 681-694.
  • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
  • GBIF (2008-continuously updated). Global Biodiversity Information Facility http://www.gbif.org/.
  • de Wilde, W.J.J.O. & Duyfjes, B.E.E. (2008). Flora of Thailand 9(4): 411-546. The Forest Herbarium, Royal Forest Department.
  • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
  • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
  • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
  • Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007). A checklist of the vascular plants of Lao PDR: 1-394. Royal Botanic Gardens, Edinburgh.
  • Trusty, J.L., Kesler, H.C. & Delgado, G.H. (2006). Vascular flora of Isla del Coco, Costa Rica Proceedings of the California Academy of Sciences, ser. 4, 57: 247-355.
  • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
  • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
  • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
  • Miller, A.G. & Morris, M. (2004). Ethnoflora of Soqotra Archipelago: 1-759. The Royal Botanic Garden, Edinburgh.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Lê, T.C. (2003). Danh l?c các loài th?c v?t Vi?t Nam 2: 1-1203. Hà N?i : Nhà xu?t b?n Nông nghi?p.
  • Grierson, A.J.C. & Long, D.G. (2001). Flora of Bhutan 2: 1-1675. Royal Botanic Gardens, Edinburgh.
  • Balick, M. J., M. H. Nee & D.E. Atha (2000). Checklist of the vascular plants of Belize Memoirs of the New York Botanical Garden 85: 1-246.
  • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
  • Dassanayake (ed.) (1997). A Revised Handbook to the Flora of Ceylon 11: 1-420. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
  • Turner, I.M. (1995). A catalogue of the Vascular Plants of Malaya Gardens' Bulletin Singapore 47(1): 1-346.
  • Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
  • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
  • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
  • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
  • Zizka, G. (1991). Flowering plants of Easter island Palmarum Hortus Francofurtensis 3: 1-108.
  • Bosser, J. & al. (eds.) (1990). Flore des Mascareignes 90-106: 1. IRD Éditions, MSIRI, RBG-Kew, Paris.
  • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
  • Troupin, G. (ed.) (1983). Flora du Rwanda 2: 1-603. Musee Royal de l'Afrique Centrale.
  • (1982). Flora of Australia 8: 1-420. Australian Government Publishing Service, Canberra.
  • Correll, D.S. & Correll, H.B. (1982). Flora of the Bahama Archipelago: 1-1692. J.Cramer, Vaduz.
  • Smith, A.C. (1981). Flora Vitiensis Nova. A new flora for Fiji (Spermatophytes only) 2: 1-810. Pacific Tropical Botanical Garden, Lawai.
  • Launert, E. (ed.) (1978). Flora Zambesiaca 4: 1-658. Royal Botanic Gardens, Kew.
  • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
  • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Fosberg, F.R. (1975). Vascular plants of Aitutaki Atoll Research Bulletin 190: 73-84.
  • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Sykes, W.R. (1970). Contributions to the flora of Niue Bulletin, New Zealand Department of Scientific and Industrial Research 200: 1-321.
  • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
  • Jeffrey, C. (1967). Flora of Tropical East Africa, Cucurbitaceae: 1-156.
  • Yuncker, T.G. (1959). Plants of Tonga Bernice P. Bishop Museum Bulletin 220: 1-283.
  • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
  • Christopherson, E. (1935). Flowering Plants of Samoa Bernice P. Bishop Museum Bulletin 128: 1-221.

Flora of Tropical East Africa

  • Keraudren, Fl. Madag. Cucurbitac: 30 (1966).
  • A. Meeuse in Bothalia 8: 51 (1962).
  • E. Lind & A. Tallantire, Some Common Flowering Plants of Uganda p. 97 (1962).
  • R. & A. Fernandes in Mem. Junta Invest. Ultram., sér. 2, 34: 73 (1962).
  • Chakrav., Monogr. Indian Cucurbitac.: 88 (1959).
  • Keay, Flora of West Tropical Africa ed. 2, 1: 212, fig. 84 (1954).
  • F. W. Andr., The Flowering Plants of the Anglo-Egyptian Sudan 1: 181, fig. 105 (1950).
  • R. O. Williams, Useful and Ornamental Plants in Zanzibar and Pemba p. 358 (1949).
  • Robyns, Flore des Spermatophytes du Parc National Albert 2: 396 (1947).
  • Exell, Cat. Vasc. Pl. S. Tomé: 184 (1944).
  • Hassib, Cucurbitac. in Egypt: 46, fig. 19 (1938).
  • Hutch. & Dalz., Flora of West Tropical Africa 1: 181, fig. 80 (1927)
  • Stuhlmann in Deutsch Ost-Afrika 10: 148 (1909).
  • Hook. f. in Flora of Tropical Africa 2: 537 (1871).
  • L., Sp. Pl.: 1009 (1753).

Catálogo de Plantas y Líquenes de Colombia
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Colombian resources for Plants made Accessible
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Flora Zambesiaca
Flora Zambesiaca
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Flora of Tropical East Africa
Flora of Tropical East Africa
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Flora of West Tropical Africa
Flora of West Tropical Africa
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Herbarium Catalogue Specimens
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Kew Backbone Distributions
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Neotropikey
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