1. Family: Malvaceae Juss.
    1. Urena Dill. ex L.

      1. This genus is accepted, and is native to Asia-Tropical, Africa, Southern America, Northern Territory, Northern America and Asia-Temperate..

    [FZ]

    Malvaceae, A. W. Exell. Flora Zambesiaca 1:2. 1961

    Habit
    Suffrutices, usually stellate-pubescent.
    Leaves
    Leaves petiolate, subentire, 3–5-palmatilobed or sinuous along the margin, palmately 3–7-nerved, with a conspicuous gland near the base of the central nerve; stipules small, linear to setaceous, usually caducous.
    Flowers
    Flowers axillary, solitary or fasciculate or crowded towards the end of the branches.
    Epicalyx
    Epicalyx campanulate, deeply 5-lobed; bracts lanceolate, acute, striate towards the base.
    Calyx
    Calyx cupulate, deeply 5-fid; lobes ovate or ovate-lanceolate, 1-nerved and keeled.
    Corolla
    Petals rose-pink or mauve, stellate-pilose outside.
    Stamens
    Staminal tube equalling the petals, the lower portion dilated and united with the petals at the base; anthers subsessile.
    Pistil
    Ovary depressed-globose, hirtellous or glabrous, of 5 free 1-ovulate carpels; styles 10, reflexed; stigmas discoid, fimbriate at the apex.
    Fruits
    Fruit subglobose, of 5 mericarps ultimately separating from the torus; mericarps trigonous, obovoid, coriaceous, indehiscent, the convex back stellately hirsute and with glochidiate spines, the flat lateral surfaces striate.
    Seeds
    Seeds obovoid-trigonous or reniform, glabrous.
    [FTEA]

    Malvaceae, Bernard Verdcourt & Geoffrey Mwachala. Pavonia, B Verdcourt; Kosteletzkya, OJ Blanchard Jr.; Gossypium, P Fryxell & B Verdcourt. Flora of Tropical East Africa. 2009

    Habit
    Perennial or annual herbs or subshrubs with stellate indumentum
    Leaves
    Leaves petiolate, subentire to deeply palmately 3–7-lobed, toothed, with often both coarse and finer teeth, palmately 3–9-veined, with a conspicuous gland near base of midrib consisting of a slit ± 1.5 mm long with thickened edges, sometimes also present on veins immediately on either side; stipules linear or setaceous, eventually deciduous
    Flowers
    Flowers solitary, fasciculate or merging into terminal pseudoracemes at the ends of the branches
    Epicalyx
    Epicalyx 5-lobed, the lobes ± rigid, narrowly triangular or lanceolate.
    Calyx
    Calyx 5-lobed, the lobes ovate to ovate-lanceolate, keeled
    Corolla
    Corolla pink or mauve, mostly with dark centre or rarely white, the petals obovate, densely stellate-pilose outside
    Stamens
    Staminal tube equalling the petals
    Ovary
    Ovary of 5 free 1-ovulate carpels.
    Style
    Styles 10, reflexed, with discoid stigmas
    Fruits
    Fruits depressed globose, of 5 indehiscent trigonous mericarps, stellate-pubescent and with dense glochidiate spines dorsally (except in endemic Australian species), striate on the flat contiguous lateral surfaces.
    Note
    Formerly usually treated as a single species but apart from the pantropical species U. lobata with numerous varieties there are certainly three other distinct species in Australia (Craven & Fryxell in Austral. Syst. Bot. 2: 455–460 (1989)). One of these and several Indian species were transferred to Pavonia by Hochreutiner in 1901 but the latter may also be better considered as Urena again. Urena and Pavonia are extremely close and the former is the earlier name. It appears there are 4–6 species of true Urena.

    Images

    Distribution

    Native to:

    Andaman Is., Angola, Assam, Bahamas, Bangladesh, Belize, Benin, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Burkina, Burundi, Cambodia, Cameroon, Cape Verde, Cayman Is., Central African Repu, Central American Pac, Chad, China South-Central, China Southeast, Christmas I., Colombia, Congo, Costa Rica, Cuba, Dominican Republic, East Himalaya, El Salvador, Ethiopia, French Guiana, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Guyana, Hainan, Haiti, Honduras, India, Ivory Coast, Jamaica, Japan, Jawa, Kenya, Laccadive Is., Laos, Leeward Is., Liberia, Madagascar, Malawi, Mali, Mauritania, Mauritius, Mexico Central, Mexico Gulf, Mexico Southeast, Mexico Southwest, Mozambique, Myanmar, Nansei-shoto, Nepal, Nicaragua, Nicobar Is., Niger, Nigeria, Northern Territory, Panamá, Paraguay, Peru, Philippines, Puerto Rico, Rodrigues, Rwanda, Réunion, Senegal, Sierra Leone, Solomon Is., Sri Lanka, Sudan, Suriname, Taiwan, Tanzania, Thailand, Tibet, Togo, Trinidad-Tobago, Uganda, Venezuela, Venezuelan Antilles, Vietnam, Windward Is., Zambia, Zaïre, Zimbabwe

    Extinct in:

    Seychelles

    Introduced into:

    Alabama, Ascension, Cook Is., Fiji, Florida, Galápagos, Lesser Sunda Is., Louisiana, Namibia, New Caledonia, Niue, Northern Provinces, Samoa, South Carolina, St.Helena, Tonga, Vanuatu, Wallis-Futuna Is., West Himalaya

    Urena Dill. ex L. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    s.coll. [Cat. no. s.n.] K001132302

    First published in Sp. Pl.: 692 (1753)

    Accepted by

    • Christenhusz, M.J.M., Fay, M.F. & Chase, M.W. (2017). Plants of the World. An illustrated encyclopedia of vascular plants: 1-792. Kew Publishing, The university of Chicago Press.

    Literature

    Flora of West Tropical Africa
    • —F.T.A. 1: 189.
    Flora Zambesiaca
    • Gen. Pl. ed. 5: 764 (1754).
    • Sp. Pl. 2: 692 (1753)
    Flora of Tropical East Africa
    • Blumea 14: 137–145 (1966)
    • E.J. 16: 361–385 (1892)
    • Gen. Pl., ed. 5: 309 (1754)
    • Sp. Pl.: 692 (1753)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0