1. Family: Apiaceae Lindl.
    1. Genus: Berula W.D.J.Koch
      1. Berula erecta (Huds.) Coville

        This species is accepted, and its native range is Temp. & Subtropical Northern Hemisphere to NE. & E. Tropical Africa.

    [FTEA]

    Umbelliferae, C.C. Townsend. Flora of Tropical East Africa. 1989

    Habit
    Weak, glabrous, erect or decumbent rhizomatous perennial, rooting at the lower nodes with clusters of white rootlets, 0.2–1 m. tall, rather sparingly branched from the base upwards.
    Stem
    Stem and branches succulent, fistular, with raised lines and alternating grooves, stems sometimes reddish about the base.
    Leaves
    Leaves simply pinnate; lowest leaves with the lamina linear-oblong in outline, ± 5–45 × 3–12 cm.; leaflets in (5–)7–11(–13) pairs, linear to broadly or oblong-lanceolate, 0.7–7.5 × 0.3–2 cm., acute to obtuse at the apex, occasionally incised at the base on the antical margin with a forwardly-directed lobule, margins with jaggedly acute to rather blunt, pale- or whitish-mucronate teeth; sheath oblong, occasionally ciliate, auriculate above, ± 0.6–4 cm.; petiole 2.5–11 cm.; upper leaves rapidly much reduced, sessile on the sheaths, leaflets often very small and sharply toothed, in 2–5 pairs.
    Inflorescences
    Umbels on 1.5–8 cm. peduncles; rays 9–16, usually unequal, 0.7–4 cm.; involucre conspicuous, of ± 5–8 linear and simple to dentate, trisect or pinnatisect, pale-margined bracts; partial umbels ± 7–22-flowered, pedicels 2–4(–6) mm.; involucel similar to the involucre but smaller, entire or sparingly denticulate, subequalling the flowers at anthesis.
    Calyx
    Sepals small but obvious, subulate.
    Corolla
    Petals white, ± 1 mm.
    Fruits
    Fruit broadly ovoid to almost round in outline, ± 2 mm., glabrous, primary ribs pale, narrow; stylopodia shortly conical, exceeded by the rather slender, ± 0.75–1 mm. styles.
    Figures
    Fig. 24.
    Habitat
    Aquatic or subaquatic, in swamps, by streams or rivers and at the edges of lakes (in one case said to be “salt”); 605–2300 m.
    Distribution
    K3 K4 T1 T2 U2 C. and S. Africawidespread in temperate Eurasia from British Is. to Himalaya, in N. America south to Mexicointroduced in Australia and elsewhere
    [FZ]

    Umbelliferae, J. F. M. Cannon. Flora Zambesiaca 4. 1978

    Habit
    Robust glabrous perennial herb up to 2·5 m. high, erect or decumbent, rhizomatous to stoloniferous, rooting at the nodes.
    Stem
    Stem hollow, terete, with coarse grooves.
    Leaves
    Leaves up to 50 cm. long, simply pinnate, with 7–12 pairs of sessile leaflets; petioles up to 12 cm. long, leaf bases narrowly sheathing. Leaflets 1–8 cm. long, coarsely and rather irregularly toothed (some of the lower leaflets sometimes showing a tendency towards the separation of leaflets of the second order). Teeth sometimes with a cartilaginous covering which may extend along the margin towards the next tooth. Upper leaves gradually reduced in size and form upwards.
    Inflorescences
    Umbels terminal and lateral on peduncles 2–12 cm. long. Bracts numerous and well developed, narrowly linear with a narrow membranous margin or linear-3-partite with obvious lateral teeth. Rays often rather irregular, 10–20; bracteoles numerous, similar to the bracts but correspondingly smaller.
    Flowers
    Flowers on fine, sometimes irregular, pedicels.
    Corolla
    Petals white, calyx teeth triangular-subulate, persistent in fruit.
    Fruits
    Fruit c. 2 mm. long, brownish at maturity, almost orbicular, slightly broader than long, depressed laterally; stylopodium 0·5 mm. long, small, conical; styles c. 1 mm. long, slightly clubbed at the apex; ribs filiform and inconspicuous at maturity.

    Images

    Distribution

    Native to:

    Afghanistan, Albania, Arizona, Austria, Baleares, Baltic States, Belarus, Belgium, British Columbia, Bulgaria, California, Central European Rus, Corse, Czechoslovakia, Denmark, East European Russia, Egypt, Eritrea, Ethiopia, France, Germany, Great Britain, Greece, Guatemala, Hungary, Illinois, Iran, Ireland, Italy, Kansas, Kazakhstan, Kenya, Kirgizstan, Krym, Lebanon-Syria, Mexico Central, Mexico Southeast, Mexico Southwest, Montana, Morocco, Nepal, Netherlands, New Mexico, North European Russi, Northwest European R, Norway, Ontario, Oregon, Pakistan, Poland, Portugal, Romania, Sardegna, Sicilia, South European Russi, Spain, Sudan, Sweden, Switzerland, Tadzhikistan, Tanzania, Texas, Turkey, Turkey-in-Europe, Turkmenistan, Uganda, Ukraine, Utah, Uzbekistan, West Himalaya, West Siberia, Wyoming, Xinjiang, Yemen, Yugoslavia

    Extinct in:

    Palestine

    Introduced into:

    Cape Provinces, Colorado, South Australia

    Berula erecta (Huds.) Coville appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Ash [859], Ethiopia 36261.000
    Wright, C. [1105], USA K001041057
    Rydberg, P.A. [722], USA K001041056
    s.coll. [s.n.] K000685625
    Sheahan, M-C. [MCS271], United Kingdom K000696386

    First published in Contr. U.S. Natl. Herb. 4: 115 (1893)

    Accepted by

    • Ackerfield, J. (2015). Flora of Colorado: 1-818. BRIT Press.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Mohlenbrock, R.H. (2014). Vascular Flora of Illinois. A Field Guide, ed. 4: 1-536. Southern Illinois University Press, Carbondale.
    • Allred, K.W. (2012). Flora Neomexicana, ed. 2, 1: 1-599. Range Science Herbarium, Las Cruces, New Mexico.
    • Danihelka, J. Chrtek, J. & Kaplan, Z. (2012). Checklist of vascular plants of the Czech Republic Preslia. Casopsi Ceské Botanické Spolecnosti 84: 647-811.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hedberg, I., Edwards, S. & Nemomissa, S. (eds.) (2003). Flora of Ethiopia and Eritrea 4(1): 1-352. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Sapir, Y., Shmida, A. & Fragman, O. (2003). Constructing Red Numbers for setting conservation priorities of endangered plant species: Israeli flora as a test case Journal for Nature Conservation 11: 1-17.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Douglas, G.W., Meidinger, D. & Penny, J.L. (2002). Rare Native Vascular Plants of British Columbia . ed. 2: 1-358. Province of British Columbia.
    • Boulos, L. (2000). Flora of Egypt 2: 1-352. Al Hadara Publishing, Cairo.
    • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
    • Govaerts, R. (1996). World Checklist of Seed Plants 2(1, 2): 1-492. MIM, Deurne.
    • Townsend, C.C. (1989). Flora of Tropical East Africa, Umbelliferae: 1-127.
    • Launert, E. (ed.) (1978). Flora Zambesiaca 4: 1-658. Royal Botanic Gardens, Kew.
    • Tutin, T.G. & al. (eds.) (1968). Flora Europaea 2: 1-469. Cambridge University Press.

    Literature

    Kew Backbone Distributions
    • Ackerfield, J. (2015). Flora of Colorado: 1-818. BRIT Press.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Allred, K.W. (2012). Flora Neomexicana, ed. 2, 1: 1-599. Range Science Herbarium, Las Cruces, New Mexico.
    • Danihelka, J. Chrtek, J. & Kaplan, Z. (2012). Checklist of vascular plants of the Czech Republic Preslia. Casopsi Ceské Botanické Spolecnosti 84: 647-811.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • Dobignard, A. & Chatelain, C. (2011). Index synonymique de la flore d'Afrique du nord 2: 1-429. Éditions des conservatoire et jardin botaniques, Genève.
    • GBIF (2008-continuously updated). Global Biodiversity Information Facility http://www.gbif.org/.
    • Hedberg, I., Edwards, S. & Nemomissa, S. (eds.) (2003). Flora of Ethiopia and Eritrea 4(1): 1-352. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Sapir, Y., Shmida, A. & Fragman, O. (2003). Constructing Red Numbers for setting conservation priorities of endangered plant species: Israeli flora as a test case Journal for Nature Conservation 11: 1-17.
    • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
    • Townsend, C.C. (1989). Flora of Tropical East Africa, Umbelliferae: 1-127.
    • Davis, P.H. (ed.) (1972). Flora of Turkey and the East Aegean Islands 4: 1-657. Edinburgh University Press, Edinburgh.
    Flora of Tropical East Africa
    • Cannon in Flora Zambesiaca 4: 600 (1978).
    • A.D.Q. Agnew, Upland Kenya Wild Flowers p. 357 (1974).
    • Cannon in Consp. Fl. Angol. 4: 349 (1970).
    • Cufod., Enumeratio Plantarum Aethiopiae Spermatophyta (Supplement in Bull. Jard. Bot. Brux.) p. 646 (1959).
    • F. W. Andr., The Flowering Plants of the Anglo-Egyptian Sudan 2: 359 (1952).
    • H. Wolff in A. Engler, Das Pflanzenreich IV. 228, heft 90: 337 (1927).
    • Cov. in Contr. U.S. Nat. Herb. 4: 115 (1893).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0