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This genus is accepted, and its native range is Cosmopolitan.


Dryopteridaceae, JP Roux, M Shaffer-Fehre, B Verdcourt. Flora of Tropical East Africa. 2007

Morphology General Habit
Plants terrestrial or epilithic
Vegetative Multiplication Rhizomes
Rhizome erect to short-decumbent, simple or sparsely branched, densely scaled
Morphology Leaves
Fronds closely spaced, axes shallowly sulcate, sulci confluent or not.
Morphology Leaves Leaf lamina
Lamina anadromous, catadromous towards apex, herbaceous to firmly herbaceous, 2–4-pinnate-pinnatifid, basal pinna pair mostly basiscopically developed, adaxially proliferous along rachis or not.
Morphology Leaves Leaf veins
Venation free, lateral veins in lobes simple, forked, or pinnately branched, vein branches end in teeth near margin, endings often slightly enlarged.
Morphology General Indumentum
Indumentum composed of unicellular glands, uniseriate pluricellular, isocytic (all cells similar) or moniliform hairs, and scales occurring on rhizome, frond axes and lamina
Morphology Reproductive morphology Sori
Sori circular, medial to inframedial on predominantly anadromous (often modified) vein branches; sporangium stalk simple, glandular and/ or haired; sori exindusiate or indusiate, indusium firmly herbaceous, deltoid to reniform, entire to erose, often glandular along margin and on surface
Morphology Reproductive morphology Spores
Spores ellipsoidal, monolete, pale to dark brown, perispore folded to form reticulate ridges or echinulate.
A genus of about 150 species with a nearly world-wide distribution.


Dryopteridaceae, JP Roux, M Shaffer-Fehre, B Verdcourt. Flora of Tropical East Africa. 2007

Morphology General Habit
Medium-sized terrestrial ferns, rhizome erect or ascending, covered in dark, entire scales
Morphology Leaves
Fronds in tufts
Morphology Leaves Stipes
Stipes densely covered in scales, adaxial groove present, grooves of axes of higher order dwindling at base
Morphology Leaves Leaf lamina
Lamina deltoid-ovate, acuminate, tripinnatifid; lowest pinna basiscopically produced, anadromous, those above catadromous.
Morphology Leaves Pinnae
Pinna bases equal, margin entire, rarely serrate.
Morphology Leaves Leaf veins
Veins dark, free, not reaching margin, apexes terminate in hydathodes
Morphology Reproductive morphology Sori
Sori round, indusiate.
A genus of five species from southern China, Taiwan, northern India, South Africa and the Madagascan region.

Native to:

Afghanistan, Alabama, Alaska, Albania, Alberta, Algeria, Altay, Amsterdam-St.Paul Is, Amur, Angola, Argentina Northwest, Arizona, Arkansas, Ascension, Assam, Austria, Azores, Baleares, Baltic States, Bangladesh, Belarus, Belgium, Bermuda, Bismarck Archipelago, Bolivia, Borneo, Brazil North, Brazil South, Brazil Southeast, Brazil West-Central, British Columbia, Bulgaria, Burundi, Buryatiya, California, Cambodia, Cameroon, Canary Is., Cape Provinces, Cape Verde, Caroline Is., Central African Repu, Central European Rus, China North-Central, China South-Central, China Southeast, Chita, Colombia, Colorado, Comoros, Connecticut, Cook Is., Corse, Costa Rica, Cuba, Cyprus, Czechoslovakia, Delaware, Denmark, District of Columbia, Dominican Republic, East European Russia, East Himalaya, Easter Is., Ecuador, El Salvador, Equatorial Guinea, Eritrea, Ethiopia, Fiji, Finland, Florida, France, Free State, Føroyar, Gabon, Galápagos, Georgia, Germany, Great Britain, Greece, Greenland, Guatemala, Guinea, Guinea-Bissau, Gulf of Guinea Is., Hainan, Haiti, Hawaii, Honduras, Hungary, Iceland, Idaho, Illinois, India, Indiana, Inner Mongolia, Iowa, Iran, Ireland, Irkutsk, Italy, Ivory Coast, Jamaica, Japan, Jawa, Kamchatka, Kansas, Kazakhstan, Kazan-retto, Kentucky, Kenya, Khabarovsk, Kirgizstan, Korea, Krasnoyarsk, Kriti, Krym, Kuril Is., KwaZulu-Natal, Labrador, Laos, Lebanon-Syria, Lesotho, Lesser Sunda Is., Liberia, Louisiana, Madagascar, Madeira, Magadan, Maine, Malawi, Malaya, Maluku, Manchuria, Manitoba, Marquesas, Maryland, Massachusetts, Mauritius, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Michigan, Minnesota, Mississippi, Missouri, Mongolia, Montana, Morocco, Mozambique, Myanmar, Nansei-shoto, Nebraska, Nepal, Netherlands, Nevada, New Brunswick, New Guinea, New Hampshire, New Jersey, New Mexico, New York, Newfoundland, Nicaragua, Nigeria, North Carolina, North Caucasus, North Dakota, North European Russi, Northern Provinces, Northwest European R, Northwest Territorie, Norway, Nova Scotia, Nunavut, Ogasawara-shoto, Ohio, Oklahoma, Ontario, Oregon, Pakistan, Palestine, Panamá, Pennsylvania, Peru, Philippines, Poland, Portugal, Primorye, Prince Edward I., Qinghai, Queensland, Québec, Rhode I., Romania, Rwanda, Réunion, Sakhalin, Samoa, Sardegna, Saskatchewan, Seychelles, Sicilia, Sierra Leone, Society Is., Solomon Is., Somalia, South Carolina, South Dakota, Spain, Sri Lanka, St.Helena, Sudan, Sulawesi, Sumatera, Swaziland, Sweden, Switzerland, Tadzhikistan, Taiwan, Tanzania, Tennessee, Texas, Thailand, Tibet, Transcaucasus, Tristan da Cunha, Tunisia, Turkey, Turkey-in-Europe, Tuva, Uganda, Ukraine, Utah, Uzbekistan, Vanuatu, Venezuela, Vermont, Vietnam, Virginia, Washington, West Himalaya, West Siberia, West Virginia, Wisconsin, Wyoming, Xinjiang, Yakutskiya, Yemen, Yugoslavia, Yukon, Zambia, Zaïre, Zimbabwe

Introduced into:

Argentina South, Chatham Is., Falkland Is.

Dryopteris Adans. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Aug 1, 2001 Etuge, M. [2062], Cameroon K000108985
Aug 1, 2001 Etuge, M. [2102], Cameroon K000108986
Kamau [621], Kenya K000696578
Kamau [622], Kenya K000696579
Comins, R.B. [s.n.], Solomon Is. Nephrodium K000636178 Unknown type material

First published in Fam. Pl. 2: 20, 551 (1763)

Accepted by

  • Fraser-Jenkins, C.R. & Kandel, D.R. (2019). Ferns and Fern-Allies of Nepal 2: 1-446. Government of Nepal, Ministry of Forests and Environment, Department of Plant Resources.
  • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.


Flora of Tropical East Africa

  • Kubitzki et al., Fam. Gen. Vasc. Pl. 1:110–112 (1990)
  • Fam. Pl. 2: 20, 551 (1763);

Flora of Tropical East Africa
Flora of Tropical East Africa

Herbarium Catalogue Specimens
'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
Digital Image © Board of Trustees, RBG Kew

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 World Checklist of Selected Plant Families.

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.