1. Family: Rubiaceae Juss.
    1. Keetia E.Phillips

      1. This genus is accepted, and its native range is Tropical & S. Africa.

    [FZ]

    Rubiaceae, D. Bridson. Flora Zambesiaca 5:2. 1998

    Habit
    Climbers or scandent shrubs; stems glabrous or frequently pubescent.
    Leaves
    Leaves paired, petiolate, not restricted to new growth at apex of branches; blades chartaceous or occasionally coriaceous; leaves subtending lateral branches often smaller and broader than main leaves; stipules interpetiolar, lanceolate to ovate or triangular at the base and acuminate to linear above, never keeled, never with white silky hairs inside.
    Flowers
    Flowers 4–6-merous, borne in pedunculate usually distinctly branched cymes; bracts and bracteoles often conspicuous.
    Calyx
    Calyx tube ellipsoid to ovoid; limb equalling or sometimes exceeding the tube in length, repand to dentate or less often lobed.
    Corolla
    Corolla white, cream or yellow; tube cylindrical, usually ± equal to lobes, sometimes shorter or rarely longer, typically with a ring of deflexed hairs inside; lobes reflexed, often thickened at apex but never apiculate.
    Stamens
    Stamens set at the throat of the corolla; filaments moderately well-developed; anthers fully or partly exserted but usually not reflexed, narrowly ovate or oblong.
    Gynoecium
    Style long slender, ± twice as long as the corolla tube; pollen presenter cylindrical, distinctly longer than wide, hollow to below the apex, bifid at apex when mature.
    Disc
    Disk puberulous to pubescent or infrequently glabrous.
    Ovary
    Ovary 2-locular, each loculus containing 1 ovule attached to upper third of the septum.
    Fruits
    Fruit a 2-seeded drupe, slightly to strongly bilobed, somewhat laterally flattened, slightly to strongly indented at apex; pyrenes woody or less often cartilaginous, usually ± ovoid with the ventral face flattened, somewhat bullate; point of attachment on the ventral face above centre or near apex; lid-like area completely or incompletely defined, either lying along ventral face above point of attachment, or across the apex, provided with a central crest, eventually dehiscent.
    Seeds
    Seeds ± ovoid, shaped at apex according to position of lid-like area in pyrene, convoluted; endosperm streaked with tanniniferous areas (resembling a ruminate endosperm, except that the testa is never vaginated) occasionally with tannin granules ± evenly dispersed or less often absent; testa thin, very finely reticulate; embryo straight with radicle erect and small cotyledons lying parallel to ventral face of the seed.
    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Climbers or scandent shrubs; stems glabrous or frequently pubescent
    Leaves
    Leaves paired, petiolate, not restricted to new growth at apex of branches; blades chartaceous or occasionally coriaceous; leaves subtending lateral branches sometimes smaller and broader than main leaves; stipules lanceolate to ovate or triangular at base and acuminate to linear above, never keeled, never with white silky hairs inside
    Flowers
    Flowers 4–6-merous, borne in pedunculate, usually distinctly branched cymes; bracts and bracteoles often conspicuous, Calyx-tube ellipsoid to ovoid; limb equalling or sometimes exceeding tube in length, repand to dentate or less often lobed
    Corolla
    Corolla white, cream or yellow; tube cylindrical, usually ± equalling lobes, sometimes shorter or rarely longer, typically with a ring of deflexed hairs inside; lobes reflexed, often thickened at apex but never apiculate
    Stamens
    Stamens set at throat of corolla; filaments moderately well developed; anthers fully or partly exserted but usually not reflexed, narrowly ovate or oblong
    Pistil
    Style long, slender, ± twice as long as the corolla-tube; stigmatic knob cylindrical, distinctly longer than wide, hollow to below apex, bifid at apex when mature (fig.131/16, p. 752); ovary 2-locular, each locule containing 1 ovule attached to upper 1/3 of the septum
    Disc
    Disk puberulous to pubescent or infrequently glabrous
    Fruits
    Fruit a 2-seeded drupe, slightly to strongly bilobed, somewhat laterally flattened, slightly to strongly indented at apex, sometimes 1-seeded by abortion and asymmetrical; pyrenes woody or less often cartilaginous, usually ± ovoid with ventral face flattened, rugulose or somewhat colliculate; point of attachment on ventral face above centre or near apex; lid-like area completely or incompletely defined, either lying along ventral face above point of attachment or across apex, provided with a central crest, eventually dehiscent around circumference (fig. 132/25, 26, p. 754)
    Seeds
    Seeds ovoid, shaped at apex according to position of lid-like area in pyrene, convoluted; endosperm streaked with tanniniferous areas (resembling a ruminate endosperm, except that the testa is never invaginated) occasionally with tannin granules ± evenly dispersed or less often absent; testa thin, very finely reticulate; embryo straight with erect radicle and small cotyledons lying parallel to ventral face of seed (fig.131/25, p. 752).

    Images

    Distribution

    Native to:

    Angola, Benin, Burkina, Burundi, Cabinda, Cameroon, Cape Provinces, Central African Repu, Congo, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Kenya, KwaZulu-Natal, Liberia, Malawi, Mali, Mozambique, Nigeria, Northern Provinces, Rwanda, Senegal, Sierra Leone, Sudan, Swaziland, Tanzania, Togo, Uganda, Zambia, Zaïre, Zimbabwe

    Keetia E.Phillips appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1994 Tchouto (Mbatchou), P. [583], Cameroon K000085364
    Jan 1, 1984 Carmichael [306], Tanzania K000319798 holotype
    Milne-Redhead, E. [5059A], Ghana 12182.000
    Milne-Redhead, E. [5059], Ghana 4687.000
    Meikle, R.D. [1392], Nigeria 5013.000
    Olorunfemi, J. [30513], Cameroon 25153.000
    Lovett, J. [3151], Tanzania K000319796
    Kayombo, C.J. [612], Tanzania K000319797

    First published in Gen. S. Afr. Fl. Pl.: 587 (1926)

    Accepted by

    • Tilney, P.M. & Van Wyk, A.E. (2009). Taxonomy of the genus Keetia (Rubiaceae - subfam. Ixoroideae - tribe Vanguerieae) in southern Africa, with notes on bacterial symbiosis as well as the structure of colleters and the 'stylar head' complex Bothalia 39: 165-175.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • Bridson in Kew Bull. 41: 965–994 (1986).
    • in Bothalia 2: 369 (1927).
    • Gen. S. African Fl. Pl. [Mem. Bot. Survey S. Afr. No. 10]: 587 (1926)
    Flora of Tropical East Africa
    • Bridson in K.B. 41: 965–994 (1986)
    • in Bothalia 2: 369 (1927)
    • in Gen. S. Afr. Fl. Pl.: 587 (1926)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0