1. Family: Rubiaceae Juss.
    1. Nauclea L.

      1. This genus is accepted, and its native range is Tropical Africa, Tropical & Subtropical Asia to N. Australia.

    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Medium-sized to large trees
    Buds
    Terminal vegetative buds strongly flattened
    Leaves
    Leaves opposite, petiolate, the nerve-axils with or without domatia; stipules ovate, obovate or elliptic, flattened, adpressed, deciduous or ± persistent
    Inflorescences
    Inflorescences terminal or terminal and lateral with unbranched axes; nodes with reduced leaves and stipules not modified into bracts which surround the young inflorescence; true peduncle glabrous or pubescent, elongating
    Flowers
    Flowers 4–5-merous, calyx glabrous or pubescent inside; limb-tube developed or not; lobes triangular or oblong, obtuse, with or without persistent appendages
    Corolla
    Corolla-tube funnel-shaped; lobes oblong, imbricate
    Stamens
    Stamens inserted in the throat, exserted; filaments short, glabrous; anthers basifixed, introrse
    Pistil
    Ovaries entirely fused to each other, 2-locular, the placentas attached to the upper third of the septum, Y-shaped with 2 short ascending arms and long basal one; ovules numerous in each locule, mostly pendulous, some erect or horizontal but these mostly aborting; style exserted; stigma spindle-shaped
    Fruits
    Fruits indehiscent, connate into a fleshy syncarp
    Seeds
    Seeds ovoid or ellipsoid, sometimes compressed, not winged.
    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Unarmed trees
    Leaves
    Leaves petiolate, large; stipules large, deciduous
    Inflorescences
    Inflorescences globose, solitary, terminal, with the flowers inserted on a fleshy ± pilose receptacle; interfloral bracteoles absent
    Calyx
    Calyx 5-lobed, the lobes free, linear-spathulate, persistent
    Corolla
    Corolla ± tubular; lobes 5, imbricate
    Stamens
    Stamen-filaments very short; anthers 2-thecous
    Pistil
    Ovaries closely contiguous but not fused, each 2-locular; locules with many ovules, the placentas attached to the middle of the septum; style slender, exserted; stigma red, narrowly conical with widened base and blunt tip, sulcate
    Fruits
    Fruiting heads globose, but individual fruits not joined, red, irregularly breaking, crowned by the persistent calyx-lobes
    Seeds
    Seeds not winged, but sometimes angular; testa reticulate; embryo straight, surrounded by albumen.
    [FZ]

    Rubiaceae, D.M.Bridson & B.Verdcourt. Flora Zambesiaca 5:3. 2003

    Habit
    Trees, unarmed.
    Buds
    Terminal vegetative buds flattened.
    Leaves
    Leaves petiolate, large; stipules large, deciduous.
    Inflorescences
    Inflorescences globose, solitary, terminal with the flowers inserted on a fleshy ± pilose receptacle.
    Calyx
    Calyx tubes closely contiguous but not fused; limb-tube ± absent; lobes 5, linear-spathulate, persistent.
    Corolla
    Corolla ± tubular; lobes 5, imbricate.
    Stamens
    Stamens with filaments very short; anthers 2-thecous.
    Gynoecium
    Style slender, exserted; pollen presenter, narrowly conical with widened base and blunt tip, sulcate, receptive areas in distal and proximal parts.
    Ovary
    Ovaries 2-locular; locules with many ovules, the placentas attached towards the top of the septum.
    Fruits
    Fruiting heads globose but individual fruits not joined, indehiscent, red, crowned by the persistent calyx lobes.
    Seeds
    Seeds not winged but sometimes angular; testa reticulate; embryo straight, surrounded by albumen.
    [FZ]

    Rubiaceae, D.M.Bridson & B.Verdcourt. Flora Zambesiaca 5:3. 2003

    Habit
    Small to medium sized trees.
    Buds
    Terminal vegetative buds flattened.
    Leaves
    Leaves opposite, petiolate, the nerve axils with or without domatia; stipules deltoid, obtuse or slightly notched at apex, ± persistent.
    Inflorescences
    Inflorescences terminal or terminal and lateral with unbranched axes, the node with reduced leaves and stipules which are not modified into bracts surrounding the young inflorescence; true peduncle glabrous or pubescent, elongating.
    Flowers
    Flowers 4–5-merous.
    Calyx
    Calyx tubes entirely fused to each other; limb-tube ± developed, densely pubescent inside, lobes persistent obtuse to triangular with or without deciduous or persistent appendages.
    Corolla
    Corolla tube funnel-shaped; lobes oblong, imbricate in the bud.
    Stamens
    Stamens inserted in the corolla tube, not exserted; anthers basifixed, introrse; filaments short, glabrous.
    Gynoecium
    Style exserted; pollen presenter spindle-shaped, the receptive areas in distal and proximal parts.
    Ovary
    Ovaries 2-locular; placentas discoidal, attached to the middle of the septum; ovules numerous in each locule, spreading in all directions.
    Fruits
    Individual fruits indehiscent, joined to form a fleshy syncarp.
    Seeds
    Seeds ovoid or ellipsoid, not winged.
    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Small to medium-sized trees
    Buds
    Terminal vegetative buds flattened
    Leaves
    Leaves opposite, petiolate, the nerve-axils with or without domatia; stipules deltoid, obtuse or slightly notched at apex, ± persistent
    Inflorescences
    Inflorescences terminal or terminal and lateral with unbranched axes, the node with reduced leaves and stipules which are not modified into bracts surrounding the young inflorescence; true peduncle glabrous or pubescent, elongating
    Flowers
    Flowers 4–5-merous, calyx densely pubescent inside, limb-tubes ± developed
    Calyx
    Calyx-lobes obtuse to triangular with or without deciduous or persistent appendages
    Corolla
    Corolla-tube funnel-shaped; lobes oblong, imbricate in the bud
    Stamens
    Stamens inserted in the tube, not exserted; filaments short glabrous; anthers basifixed, introrse
    Pistil
    Ovaries entirely fused to each other, 2-locular; placentas ± discoidal, attached to the middle of the septum; ovules numerous in each locule, spreading in all directions; style exserted; stigma spindle-shaped
    Fruits
    Individual fruits indehiscent, joined to form a fleshy syncarp
    Seeds
    Seeds ovoid or ellipsoid, not winged.

    Images

    Distribution

    Native to:

    Angola, Assam, Bangladesh, Benin, Bismarck Archipelago, Borneo, Burkina, Cabinda, Cambodia, Cameroon, Central African Repu, Chad, China Southeast, Congo, Equatorial Guinea, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Hainan, Ivory Coast, Jawa, Kenya, Laos, Lesser Sunda Is., Liberia, Malawi, Malaya, Mali, Maluku, Mauritania, Mozambique, Myanmar, New Guinea, Nigeria, Northern Territory, Philippines, Queensland, Senegal, Sierra Leone, Solomon Is., Sri Lanka, Sudan, Sulawesi, Sumatera, Tanzania, Thailand, Togo, Uganda, Vietnam, Zambia, Zaïre

    Introduced into:

    Fiji

    Nauclea L. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Sierra Leone 36204.000

    First published in Sp. Pl. ed. 2: 243 (1762)

    Accepted by

    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora of West Tropical Africa
    • Merrill in J. Wash. Acad. Sci. 5: 530 (1915).
    • Sp. Pl., ed. 2, 1: 243 (1762)
    Flora of Tropical East Africa
    • Ridsd. in Blumea 24: 325 (1978)
    • Ridsd. in Blumea 23: 184–186 (1976)
    • Ridsd. in Blumea 22: 547 (1975)
    • Merr. in Journ. Wash. Acad. Sci. 5: 534 (1915)
    • Sp. Pl., ed. 2, 1: 243 (1762)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0