1. Family: Phyllanthaceae Martinov
    1. Genus: Cleistanthus Hook.f. ex Planch.
      1. Cleistanthus schlechteri (Pax) Hutch.

        This species is accepted, and its native range is Kenya to KwaZulu-Natal.

    [FZ]

    Euphorbiaceae, A. Radcliffe-Smith. Flora Zambesiaca 9:4. 1996

    Habit
    A many-stemmed shrub or large tree up to c. 20 m tall with a rounded crown and drooping ultimate branchlets.
    Bole
    Bole to 3.5 m high, 4 dm in diameter.
    Bark
    Bark on trunk dark grey or reddish-brown to blackish, rough, reticulate-striate, fibrous, paler on the branches.
    Wood
    Wood hard.
    Twigs
    Twigs grey, lenticellate.
    Shoots
    Young shoots and petioles pubescent to subglabrous.
    Petiole
    Petioles 2–7 mm long.
    Stipules
    Stipules 4–5 × 1–2 mm, narrowly lanceolate to falcate-lanceolate, sparingly pubescent, membranaceous, fugacious.
    Leaf lamina
    Leaf blades 1–9(12) × 0.5–4.5(5.5) cm, elliptic-ovate to elliptic-oblong, obtuse to obtusely acuminate, sometimes emarginate, cuneate-rounded to shallowly cordate at the base, epipetiolar, firmly chartaceous, very sparingly appressed-pubescent along the midrib beneath and otherwise glabrous, more rarely evenly patent-pubescent along the midrib beneath, dark green and shiny above, paler and duller beneath, drying greenish or greyish-green; lateral nerves in 5–8 pairs, brochidodromous, scarcely to slightly prominent above and beneath, bearing acarodomatia in their axils in larger leaves, tertiary nerves reticulate.
    Inflorescences
    Inflorescences up to 12-flowered, either all male or male with 1(2) female flowers; axes less than 1 cm long, appressed-pubescent; bracts c. 1 × 1 mm, ± ovate, caducous.
    Flowers
    Flowers fragrant. Female flowers: pedicels extending to 3 cm long in fruit, pubescent, reddish; buds 5 mm long, ovoid-conic; sepals and petals ± as in the male; disk 2–2.5 mm in diameter, otherwise as in the male; ovary 3 mm in diameter, globose, glabrous or else densely appressed golden-pubescent; styles 3, 2 mm long, united for 0.5 mm, once bifid, glabrous. Male flowers: pedicels 6–11(15) mm long, sparingly puberulous; buds 3–5 mm long, ellipsoid, rounded at the base; sepals 5–6 × 1–1.5 mm, linear, acute, cucullate, sparingly puberulous without, glabrous within, yellowish-green; petals 1–1.3 mm long, subulate; disk 1.3 mm in diameter, shallowly cupular, slightly pentagonal, glabrous, pink; staminal column 1.5 mm high; anthers 2.5–3 mm long, cream-coloured; pistillode 1 mm high, tripartite, glabrous.
    Male
    Male flowers: pedicels 6–11(15) mm long, sparingly puberulous; buds 3–5 mm long, ellipsoid, rounded at the base; sepals 5–6 × 1–1.5 mm, linear, acute, cucullate, sparingly puberulous without, glabrous within, yellowish-green; petals 1–1.3 mm long, subulate; disk 1.3 mm in diameter, shallowly cupular, slightly pentagonal, glabrous, pink; staminal column 1.5 mm high; anthers 2.5–3 mm long, cream-coloured; pistillode 1 mm high, tripartite, glabrous.
    Female
    Female flowers: pedicels extending to 3 cm long in fruit, pubescent, reddish; buds 5 mm long, ovoid-conic; sepals and petals ± as in the male; disk 2–2.5 mm in diameter, otherwise as in the male; ovary 3 mm in diameter, globose, glabrous or else densely appressed golden-pubescent; styles 3, 2 mm long, united for 0.5 mm, once bifid, glabrous.
    Fruits
    Fruit 7–10 × 9–15 mm, strongly 3-lobed or rarely 4-lobed, ± smooth to slightly rugulose, glabrous or evenly appressed golden-pubescent, greenish to light brown with 6 reddish lines at first, brownish-black when ripe.
    Seeds
    Seeds 5 × 4.5 mm, subglobose, smooth, shiny, brownish-pink to dark brown or blackish.
    [FTEA]

    Euphorbiaceae, A. R.-Smith. Flora of Tropical East Africa. 1987

    Habit
    A shrub or tree up to 23 m. with spreading branches, monoecious.
    Bark
    Bark grey or brown with narrow longitudinal fissures.
    Twigs
    Twigs grey, evenly minutely lenticellate.
    Shoots
    Young shoots and petioles evenly to sparingly pubescent.
    Leaves
    Petioles 4–7 mm. long; leaf-blades elliptic-ovate, elliptic-lanceolate or elliptic-oblong, 3–9 cm. long, 1–4 cm. wide, obtuse or emarginate, rarely perceptibly obtusely acuminate, rounded or shallowly cordate, the two halves meeting above the petiole, firmly chartaceous, lateral nerves 5–8 pairs, brochidodromous, slightly prominent above and beneath, tertiary nerves reticulate, scarcely prominent, very sparingly pubescent along the midrib above and beneath, otherwise quite glabrous, bright green, drying greenish or grey-green.
    Stipules
    Stipules ovate-lanceolate, 5 mm. long, 2 mm. wide, acute, sparingly pubescent, membranaceous, translucent, very readily caducous.
    Inflorescences
    Inflorescences axillary, pseudoracemose, 4–12-flowered, mostly ♂, usually with one ♀ at the base; axis condensed, 2–8 mm. long, densely yellow-pubescent; bracts lanceolate, 1 mm. long.
    Flowers
    Flowers fragrant. Female flowers:pedicels 0.5–1.5 cm. long, extending to 2.5 cm. in fruit, sparingly pubescent; buds ovoid; sepals and petals ± as in the ♂ flowers; disc annular, 2 mm. diameter, ± entire, glabrous; ovary globose, 3 mm.  long and wide, glabrous or densely adpressed golden-pubescent; styles 3, free, 2 mm. long, bifid, glabrous, cream. Male flowers:pedicels 6–11 mm. long, sparingly pubescent; buds ellipsoid, rounded at the base; sepals narrowly oblong-lanceolate, 4–6 mm. long, 1–1.5 mm. wide, acutely acuminate, cucullate, sparingly pubescent without, glabrous within, yellow or yellowish green; petals 1 mm. long, subulate; disc shallowly cupular, 1.5 mm. diameter, ± entire, glabrous; staminal column 1 mm. high; anthers 3 mm. long, cream; pistillode tripartite, 1 mm. high, glabrous.
    Male
    Male flowers:pedicels 6–11 mm. long, sparingly pubescent; buds ellipsoid, rounded at the base; sepals narrowly oblong-lanceolate, 4–6 mm. long, 1–1.5 mm. wide, acutely acuminate, cucullate, sparingly pubescent without, glabrous within, yellow or yellowish green; petals 1 mm. long, subulate; disc shallowly cupular, 1.5 mm. diameter, ± entire, glabrous; staminal column 1 mm. high; anthers 3 mm. long, cream; pistillode tripartite, 1 mm. high, glabrous.
    Female
    Female flowers:pedicels 0.5–1.5 cm. long, extending to 2.5 cm. in fruit, sparingly pubescent; buds ovoid; sepals and petals ± as in the ♂ flowers; disc annular, 2 mm. diameter, ± entire, glabrous; ovary globose, 3 mm.  long and wide, glabrous or densely adpressed golden-pubescent; styles 3, free, 2 mm. long, bifid, glabrous, cream.
    Fruits
    Fruit trilobate, 7–8 mm. long, 9–13 mm. diameter, glabrous or evenly adpressed golden-pubescent.
    Seeds
    Seeds subglobose, 5 mm. long, 4.5 mm. wide, smooth, shiny, dark chocolate-brown or blackish.

    Images

    Distribution

    Native to:

    Kenya, KwaZulu-Natal, Malawi, Mozambique, Northern Provinces, Tanzania, Zimbabwe

    Cleistanthus schlechteri (Pax) Hutch. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Aug 1, 1960 Schlechter [11524], Mozambique K000406060

    First published in W.H.Harvey & auct. suc. (eds.), Fl. Cap. 5(2): 382 (1915)

    Accepted by

    • Calane da Silva, M., Izdine, S. & Amuse, A.B. (2004). A Preliminary Checklist of the Vascular Plants of Mozambique: 1-184. SABONET, Pretoria.
    • Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.

    Literature

    Flora of Tropical East Africa
    • J. Léon. in Bulletin du Jardin Botanique de l'État a Bruxelles 30: 422 (1960).
    • A. Engler & O. Drude, Die Vegetation Der Erde, IX, Pflanzenwelt Afrikas 3(2): 41 (1921).
    • Hutch. in Fl. Cap. 5(2): 382 (1915).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0