1. Family: Apiaceae Lindl.
    1. Berula W.D.J.Koch

      1. This genus is accepted, and its native range is Africa, Temp. Northern Hemisphere to Mexico.

    [FZ]

    Umbelliferae, J. F. M. Cannon. Flora Zambesiaca 4. 1978

    Habit
    Robust glabrous herbs with pinnate leaves and hollow stems.
    Flowers
    Flowers in compound umbels with well-developed bracts and bracteoles.
    Calyx
    Sepals subulate-triangular, quite conspicuous.
    Corolla
    Petals broadly obcordate, with an inflexed point at the apex.
    Fruits
    Fruit suborbicular, almost didymous, laterally depressed and somewhat constricted at the commissure; ribs poorly developed and not conspicuous, lateral ribs not marginal. Mericarps subpentagonal in section.
    Carpophore
    Carpophore adnate to the mericarps.
    Vittae
    Vittae deeply embedded in the mesocarp.
    [FTEA]

    Umbelliferae, C.C. Townsend. Flora of Tropical East Africa. 1989

    Leaves
    Leaves simply pinnate with numerous toothed leaflets, teeth pale-mucronate
    Inflorescences
    Umbels numerous, compound, pedunculate, leaf-opposed; involucre and involucel conspicuous, of several linear and simple to toothed or dissected bracts and bracteoles
    Flowers
    Flowers hermaphrodite
    Calyx
    Calyx-teeth small but distinct
    Corolla
    Petals white, emarginate above with an incurved acute lobule, glabrous
    Fruits
    Fruit glabrous, slighdy laterally compressed, broadly ovoid to subglobose, mericarps narrowed to the commissure, subdidymous; primary ribs pale and conspicuous but scarcely at all raised, filiform; vittae deeply immersed beneath the thick spongy pericarp, not visible in surface view, in section quite large and abutting on to each other, forming a complete ring usually of ± 12–13 around the endosperm; stylopodia shortly conical, styles rather slender; carpophore adnate to the mericarps
    Seeds
    Endosperm almost round in section to very obtusely pentagonal.
    Habit
    Aquatic perennial
    [FZ]

    Umbelliferae, J. F. M. Cannon. Flora Zambesiaca 4. 1978

    Habit
    Perennial herbs with pinnate leaves; leaflets imbricated when young and bearing spinous cartilaginous teeth.
    Inflorescences
    Umbels compound; bracts and bracteoles numerous and well developed.
    Corolla
    Petals suborbicular to broadly obcordate, apex emarginate with an inflexed point.
    Calyx
    Calyx teeth obsolete.
    Stylopodium
    Stylopodium well developed, conical with a thickened, undulate margin; styles short, stiff, erect, slightly divergent; stigmatic surfaces slightly thickened.
    Fruits
    Fruit ovoid, slightly laterally compressed; ribs filiform but quite distinct. Mericarps pentagonal in section; commissural face flat.
    Vittae
    Vittae 1 in each interval and 2 in the commissural face.
    Seeds
    Commissural face of the seed flat.
    [FTEA]

    Umbelliferae, C.C. Townsend. Flora of Tropical East Africa. 1989

    Habit
    Perennial herbs with a rhizomatous rootstock
    Leaves
    Leaves simply pinnate with numerous, toothed, cartilaginous-mucronate leaflets
    Inflorescences
    Umbels several, compound, pedunculate; involucre and involucel conspicuous, bracts commonly ± divided, bracteoles entire
    Flowers
    Flowers polygamous
    Calyx
    Calyx-teeth inconspicuous or absent
    Corolla
    Petals emarginate above with an incurved, rather broad apical lobule, glabrous
    Fruits
    Fruit glabrous, broadly ovoid, somewhat laterally compressed with the primary ribs narrow but pale and prominent, commissure narrow at maturity; vittae solitary in the valleculae, 2 on the commissure; stylopodia relatively large, styles rather short and stout; carpophore bifid to the base
    Seeds
    Endosperm distinctly pentagonal, the surface slightly sulcate beneath the vittae.

    Images

    Distribution

    Native to:

    Afghanistan, Albania, Angola, Arizona, Austria, Baleares, Baltic States, Belarus, Belgium, British Columbia, Bulgaria, Burundi, California, Cameroon, Cape Provinces, Central African Repu, Central European Rus, Corse, Czechoslovakia, Denmark, East European Russia, Egypt, Eritrea, Ethiopia, France, Free State, Germany, Great Britain, Greece, Guatemala, Hungary, Illinois, Iran, Ireland, Italy, Kansas, Kazakhstan, Kenya, Kirgizstan, Krym, KwaZulu-Natal, Lebanon-Syria, Lesotho, Malawi, Mexico Central, Mexico Southeast, Mexico Southwest, Montana, Morocco, Mozambique, Namibia, Nepal, Netherlands, New Mexico, North European Russi, Northern Provinces, Northwest European R, Norway, Ontario, Oregon, Pakistan, Poland, Portugal, Romania, Sardegna, Sicilia, South European Russi, Spain, St.Helena, Sudan, Swaziland, Sweden, Switzerland, Tadzhikistan, Tanzania, Texas, Turkmenistan, Uganda, Ukraine, Utah, Uzbekistan, West Himalaya, West Siberia, Wyoming, Xinjiang, Yemen, Yugoslavia, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Colorado, South Australia

    Berula W.D.J.Koch appears in other Kew resources:

    First published in J.C.Röhling, Deutschl. Fl., ed. 3, 2: 433 (1826)

    Accepted by

    • Govaerts, R. (1996). World Checklist of Seed Plants 2(1, 2): 1-492. MIM, Deurne.

    Literature

    Flora Zambesiaca
    • in Röhl., Deutschl. Fl. 2: 25 (1826).
    Flora of Tropical East Africa
    • in Röhl., Deutsch. Fl. 2: 25, 433 (1826)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0