1. Family: Fabaceae Lindl.
    1. Genus: Mucuna Adans.
      1. Mucuna pruriens (L.) DC.

        This species is accepted, and its native range is Tropical & Subtropical Old World. It is used to treat unspecified medicinal disorders.

    [UPB]

    The Useful Plants of Boyacá project

    Distribution
    Naturalised in Colombia.
    Habit
    Liana, creeper.
    Ecology
    Alt. 80 - 1800 m.
    [FZ]

    Leguminosae, B. Mackinder, R. Pasquet, R. Polhill and B. Verdcourt. Flora Zambesiaca 3:5. 2001

    Habit
    Climbing herb 1–4.5(6) m long, usually annual or a short-lived perennial but also stated to climb to the tops of tall trees.
    Stem
    Stems densely hairy at first, later glabrous.
    Leaflets
    Leaflets rather thin, 4.8–19 × 3.4–16.8 cm, obovate, elliptic, rhombic or ovate, the laterals very oblique, acute or acuminate or rarely rounded at the apex, rounded at the base, sparsely to densely appressed grey or silvery hairy on both surfaces, not blackening on drying; petiole 2–40 cm long; rhachis 1–5 cm long; petiolules 3–6 mm long.
    Inflorescences
    Inflorescences silvery pubescent; rhachis 2–29 cm long; peduncle 2–18 cm long; flowers 2–3 per node, on pedicels 1.5–6 mm long; bracts 10–15 × 6 mm; bracteoles 10 × 2 mm, very deciduous.
    Calyx
    Calyx appressed silvery pubescent; tube 5–7 mm long; lobes 3–9 mm long, the lower linear-lanceolate to triangular, the upper pair joined to form a triangular lip.
    Corolla
    Standard blackish-purple to pale lilac or white, 1.7–2.2 × 1.4–1.5 cm, ovate, investing wings and not becoming erect; wings pale lilac to purple, 3–3.5 cm long; keel white, tinged mauve, 3.2–3.7 cm long, apical beak recurved and horny; staminal tube white.
    Fruits
    Pods 4–9 × 1–1.5 cm (sometimes 2 cm in cultivated forms), oblong, often S-shaped, densely covered with brown or reddish-orange irritant bristly hairs which sometimes appear black in the mass, longitudinally ribbed under the hairs; in some cultivated forms the pods are glabrescent or velvety hairy but lack the bristles.
    Seeds
    Seeds pinkish-brown speckled black or almost entirely black (or white to black in cultivars), 10–19 × 7.5–13 × 4–6.5 mm, oblong-ellipsoid, compressed; hilum c. 4 mm long, oblong; rim aril cream, somewhat wrinkled and with a scale-like extension.
    [CPLC]

    Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

    Distribution
    Naturalizada en Colombia; Alt. 80 - 1800 m.; Andes, Orinoquia, Pacífico, Valle del Cauca, Valle del Magdalena.
    Habit
    Trepadora, liana
    Conservation
    No Evaluada
    [FTEA]

    Leguminosae, J. B. Gillett, R. M. Polhill & B. Verdcourt. Flora of Tropical East Africa. 1971

    Habit
    Climbing herb 2–3 m. long, usually annual or a short-lived perennial, but also stated to climb to the tops of tall trees.
    Stem
    Stems densely hairy at first, later glabrous.
    Leaves
    Leaflets rather thin, obovate, elliptic, rhomboid or ovate, the laterals very oblique, 4·8–19 cm. long, 3·4–16·8 cm. wide, acute or acuminate or rarely rounded at the apex, rounded at the base, sparsely to densely adpressed grey or silvery hairy on both surfaces, not blackening on drying; petiole 2–40 cm. long; rhachis 1–5 cm. long; petiolules 3–6 mm. long.
    Inflorescences
    Inflorescences silvery pubescent; rhachis 2–29 cm. long; peduncle 2–18 cm. long; flowers 2–3 per node, on pedicels 1·5–6 mm. long; bracts 1–1·5 cm. long, 6 mm. wide; bracteoles very deciduous, 1 cm. long, 2 mm. wide.
    Calyx
    Calyx adpressed silvery pubescent; tube 5–7 mm. long; lobes 3–9 mm. long, the lower linear-lanceolate to triangular, the upper pair joined to form a triangular lip.
    Corolla
    Standard blackish-purple to pale lilac or white, ovate, 1·7–2·2 cm. long, 1·4–1·5 cm. wide; wings pale lilac to purple, 3–3·5 cm. long; keel white, tinged mauve, 3·2–3·7 cm. long, apical beak recurved and horny; staminal tube white.
    Fruits
    Pods oblong, often S-shaped, 4–9 cm. long, 1–1·5 cm. wide (sometimes 2 cm. in cultivated forms), densely covered with brown or reddish-orange irritant bristly hairs which sometimes appear black in the mass, longitudinally ribbed under the hairs; in some cultivated forms the pods are glabrescent or velvety hairy but lack the bristles.
    Seeds
    Seeds pinkish-brown speckled black or almost entirely black (or white to black in cultivars), oblong-ellipsoid, compressed, longest dimension 1–1·9 cm., shorter dimension 7·5–13 mm., thickness 4–6·5 mm.; hilum oblong, ± 4 mm. long; rim-aril cream, somewhat wrinkled and with a scale-like extension.
    [ILDIS]

    International Legume Database and Information Service

    Conservation
    Not Threatened
    Ecology
    Africa: Sudanian wooded grassland, Lake Victoria regional mosaic; forest., Lake Victoria regional mosaic; anthropic landscapes., Lake Victoria regional mosaic; bushland and thicket, Somalia-Masai wooded grasslands., Somalia-Masai bushland and thicket, Gui
    Habit
    Annual/Perennial, Climbing, Herb/Shrub
    Vernacular
    Achariya, Achariya-pala, Aga, Agy, Ahagy, Bhainswalibel, Buchariwa, Bu-chariya, Buffalo Bean, Chunao-avarai, Cow Itch, Cowage Velvetbean, Cowitch, Cow-itch, Fogarate, Ginipus-wel, Gratey, Hell Fire Bean, Itchy Bean, Phandatullai, Pica Pica, Poil A Gratter
    [UPB]
    Materials
    Materials (State of the World's Plants 2016).
    Unspecified Materials Chemicals
    Materials (State of the World's Plants 2016).
    Unspecified Medicinal Disorders
    Medicinal (State of the World's Plants 2016, Instituto Humboldt 2014).
    [ILDIS]
    Use
    Domestic, Environmental, Food and Drink, Forage, Medicine, Toxins, Weed

    Images

    Distribution

    Native to:

    Andaman Is., Angola, Assam, Bangladesh, Benin, Burkina, Burundi, Cambodia, Cameroon, Cape Verde, Central African Repu, Chad, China South-Central, Christmas I., Comoros, Congo, East Himalaya, Equatorial Guinea, Gabon, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Hainan, India, Ivory Coast, Kenya, KwaZulu-Natal, Laccadive Is., Liberia, Madagascar, Malawi, Malaya, Maluku, Mozambique, Myanmar, Nepal, Nicobar Is., Nigeria, Northern Provinces, Pakistan, Philippines, Senegal, Sierra Leone, Solomon Is., Sri Lanka, Sudan, Tanzania, Thailand, Togo, Uganda, Vietnam, West Himalaya, Yemen, Zambia, Zaïre, Zimbabwe

    Introduced into:

    Bahamas, Belize, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, China Southeast, Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, El Salvador, Ethiopia, Fiji, Guatemala, Haiti, Honduras, Jamaica, Jawa, Leeward Is., Lesser Sunda Is., Mauritius, Mexico Central, Mexico Gulf, Mexico Southeast, Mexico Southwest, New Caledonia, Nicaragua, Northern Territory, Panamá, Peru, Puerto Rico, Queensland, Réunion, Southwest Caribbean, Suriname, Taiwan, Transcaucasus, Trinidad-Tobago, Venezuela, Windward Is.

    Common Names

    Spanish
    Ojo de venado, pica pica, guisante negro, chiporro.

    Mucuna pruriens (L.) DC. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Dec 22, 2005 Garcia, F.C.P. [786], Brazil K000931379
    Nov 22, 2005 Garcia, F.C.P. [785], Brazil K000931380
    Nov 22, 2005 Garcia, F.C.P. [783], Brazil K000931375
    Nov 22, 2005 Garcia, F.C.P. [793], Brazil K000931376
    Nov 22, 2005 Garcia, F.C.P. [791], Brazil K000931377
    Nov 22, 2005 Garcia, F.C.P. [784], Brazil K000931378
    Apr 7, 2003 Hatschbach, G.G. [70801], Brazil K000931374
    Jan 1, 1995 von Rege, I. [43], Cameroon K000437643
    Jul 1, 1992 Hatschbach, G. [56954], Bahia K000206228
    Tweedie, E.M. [1038], Kenya 9279.000
    Santos, T.S. [3613], Brazil K000931373
    Queiroz, L.P. [4122], Bahia K000206671
    Queiroz, L.P. [5023], Bahia K000207268
    Wallich, N. [5617], India K000797549
    Wallich, N. [Cat. no. 5616] K001121468
    Wallich, N. [Cat. no. 5616] K001121469
    Wallich, N. [Cat. no. 5616] K001121470
    Wallich, N. [Cat. no. 5616] K001121471
    Silva, M.A. [2541], Brazil K000931372
    Porter, T.A. [s.n.], Pernambuco K000206229
    Gomez, W. [Cat. no. 5616] K001121466
    s.coll. [Cat. no. 5616] K001121461
    s.coll. [Cat. no. 5616] K001121462
    s.coll. [Cat. no. 5616] K001121463
    s.coll. [Cat. no. 5616] K001121464
    s.coll. [Cat. no. 5616] K001121465
    s.coll. [Cat. no. 5616] K001121467
    s.coll. [Cat. no. 5616] K001121472

    First published in Prodr. 2: 405 (1825)

    Accepted by

    • De Moura, T.M., Lewis, G.P., Mansano, V.F. & Tozzi, A.M.G.A. (2018). A revision of the neotropical Mucuna species (Leguminosae-Papilionoideae) Phytotaxa 337: 1-65.
    • Forzza, R.C., Zappi, D. & Souza, V.C. (2016-continuously updated). Flora do Brasil 2020 em construção http://reflora.jbrj.gov.br/reflora/listaBrasil/ConsultaPublicaUC/ResultadoDaConsultaNovaConsulta.do.
    • Wiriadinata, H., Ohashi, H. & Adema, F. (2016). Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna Blumea 61: 90-124.
    • Villaseñor, J.L. (2016). Checklist of the native vascular plants of Mexico Revista Mexicana de Biodiversidad 87: 559-902.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Lepschi, B. & Monro, A. (Project Coordinators) (2014). Australian Plant Census (APC) Council of Heads of Australian Herbaria. http://www.anbg.gov.au/chah/apc/index.html.
    • Sikarwar, R.L.S. (2014). Angiosperm diversity assessment of Chitrakootthe legendary place of Vindhyan range, India Journal of Economic and Taxonomic Botany 38: 563-619.
    • Moura, T.M., Lewis, G.P., Mansano, V.F. & Tozzi, A.M.G.A. (2014). Taxonomic studies in Mucuna Adans. (Leguminosae - Papilionoideae) from Peru Systematic Botany 39: 884-896.
    • Choudhary, R.K., Srivastava, R.C., Das, A.K. & Lee, J. (2012). Floristic diversity assessment and vegetation analysis of Upper Siang district of eastern Himalaya in North East India Korean Journal of Plant Taxonomy 42: 222-246.
    • Meena, S.L. (2012). A checklist of the vascular plants of Banaskantha district, Gujarat, India Nelumbo 54: 39-91.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • de la Estrella, M., Cabezas, F.J., Aedo, C. & Velayos, M. (2010). The Papilionoideae (Leguminosae) of Equatorial Guinea (Annobón, Bioko and Río Muni) Folia Geobotanica 45: 1-57.
    • Araújo, D. & Alves, M. (2010). Climbing plants of a fragmented area of lowland Atlantic forest, Igarassu, Pernambuco (northeastern Brazil) Phytotaxa 8: 1-24.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
    • Wu, Z. & Raven, P. (eds.) in Flora of China Editorial Committee (2010). Flora of China 10: 1-642. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
    • Sarmah, K.K. & Borthakur, S.K. (2009). A checklist of angiospermic plants of Manas national park in Assam, India Pleione 3: 190-200.
    • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Wilmot-Dear, C.M. (2008). Mucuna Adans. (Leguminosae) in Thailand Thai Forest Bulletin (Botany) 36: 114-139.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Lock, J.M. & Ford, C.S. (2004). Legumes of Malesia a Check-List: 1-295. Royal Botanic Gardens, Kew.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Kumar, S. & Sane, P.V. (2003). Legumes of South Asia. A Checklist: 1-536. Royal Botanic Gardens, Kew.
    • Du Puy, D.J., Labat, N.-N., Rabevohitra, R., Villiers, J.-F., Bosser, J. & Moat, J. (2002). The Leguminosae of Madagascar: 1-737. Royal Botanic Gardens, Kew.
    • Mackinder, B., Pasquet, R., Polhill, R. & Verdcourt, B. (2001). Flora Zambesiaca 3(5): 1-261. Royal Botanic Gardens, Kew.
    • Stevens, W.D., Ulloa U., C., Pool, A. & Montiel, O.M. (2001). Flora de Nicaragua Monographs in Systematic Botany from the Missouri Botanical Garden 85: i-xlii, 1-2666. Missouri Botanical Garden.
    • Balick, M.J., Nee, M.H. & Atha, D.E. (2000). Checklist of the Vascular Plants of Belize with Common Names an Uses: 1-246. New Yourk Botanic Garden Press, New York.
    • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
    • Jørgensen, P.M. & León-Yánes, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador: 1-1181. Missouri Botanical Garden Press, St. Louis.
    • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
    • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
    • Gonzalez, F., Nelson Diaz, J. & Lowry, P. (1995). Flora Illustrada de San Andrés y Providencia: 1-281. Universidad Nacional de Colombia, Instituto de Ciencias Naturales, Colombia.
    • Lock, J.M. & Heald, J. (1994). Legumes of Indo-China a checck-list: 1-164. Royal Botanic Gardens, Kew.
    • George, A.S., Orchard, A.E. & Hewson, H.J. (eds.) (1993). Oceanic islands 2 Flora of Australia 50: 1-606. Australian Government Publishing Service, Canberra.
    • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
    • Polhill, R.M. (1990). Flore des Mascareignes 80: 1-235. IRD Éditions, MSIRI, RBG-Kew, Paris.
    • Lock, J.M. (1989). Legumes of Africa a check-list: 1-619. Royal Botanic Gardens, Kew.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 1: 1-114. ORSTROM, Bangui.
    • Berhaut, J. (1976). Flore illustrée du Sénégal 5: 1-658. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
    • Standley, P.C. & Steyermark, J.A. (1946). Flora of Guatemala Fieldiana Botany New Series 24(5): 1-502. Field Museum of Natural History.

    Literature

    Useful Plants of Boyacá Project
    • Kew's Economic Botany collection in The State of the World’s Plants Report–2016. (2016). Royal Botanic Gardens, Kew https://stateoftheworldsplants.org/2016/
    • Instituto de Investigación de Recursos Biológicos Alexander von Humbodlt (2014). Plantas alimenticias y medicinales nativas de Colombia. 2567 registros, aportados por: Castellanos, C. (Contacto del recurso), Valderrama, N. (Creador del recurso, Autor), Castro, C. (Proveedor de metadatos), Bernal, Y. (Autor), García, N. (Autor). Versión 11.0. http://i2d.humboldt.org.co/ceiba/resource.do?r=ls_colombia_magnoliophyta_2014
    Kew Backbone Distributions
    • Forzza, R.C., Zappi, D. & Souza, V.C. (2016-continuously updated). Flora do Brasil 2020 em construção http://reflora.jbrj.gov.br/reflora/listaBrasil/ConsultaPublicaUC/ResultadoDaConsultaNovaConsulta.do.
    • Villaseñor, J.L. (2016). Checklist of the native vascular plants of Mexico Revista Mexicana de Biodiversidad 87: 559-902.
    • Ibarra-Manríquez, G. & al. (2015). Lianas of Mexico Botanical Sciences 93: 365-417.
    • Lepschi, B. & Monro, A. (Project Coordinators) (2014). Australian Plant Census (APC) Council of Heads of Australian Herbaria. http://www.anbg.gov.au/chah/apc/index.html.
    • Moura, T.M., Lewis, G.P., Mansano, V.F. & Tozzi, A.M.G.A. (2014). Taxonomic studies in Mucuna Adans. (Leguminosae - Papilionoideae) from Peru Systematic Botany 39: 884-896.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
    • Moura, T.M. & al. (2013). A taxonomic revision of Mucuna (Fabaceae: Papilionoideae: Phaseoleae) in Brazil Systematic Botany 38: 631-637.
    • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
    • Choudhary, R.K., Srivastava, R.C., Das, A.K. & Lee, J. (2012). Floristic diversity assessment and vegetation analysis of Upper Siang district of eastern Himalaya in North East India Korean Journal of Plant Taxonomy 42: 222-246.
    • Meena, S.L. (2012). A checklist of the vascular plants of Banaskantha district, Gujarat, India Nelumbo 54: 39-91.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • de la Estrella, M., Cabezas, F.J., Aedo, C. & Velayos, M. (2010). The Papilionoideae (Leguminosae) of Equatorial Guinea (Annobón, Bioko and Río Muni) Folia Geobotanica 45: 1-57.
    • Hammel, B.E., Grayum, M.H., Herrera & C. & Zamora, N. (eds.) (2010). Manual de plantas de Costa Rica volumen V. Dicotiledóneas (Clusiaceae-Gunneraceae) Monographs in Systematic Botany from the Missouri Botanical Garden 119: 1-970. Missouri Botanical Garden.
    • Araújo, D. & Alves, M. (2010). Climbing plants of a fragmented area of lowland Atlantic forest, Igarassu, Pernambuco (northeastern Brazil) Phytotaxa 8: 1-24.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania 30: 1-307.
    • Wu, Z. & Raven, P. (eds.) in Flora of China Editorial Committee (2010). Flora of China 10: 1-642. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
    • Sarmah, K.K. & Borthakur, S.K. (2009). A checklist of angiospermic plants of Manas national park in Assam, India Pleione 3: 190-200.
    • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Wilmot-Dear, C.M. (2008). Mucuna Adans. (Leguminosae) in Thailand Thai Forest Bulletin (Botany) 36: 114-139.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Lock, J.M. & Ford, C.S. (2004). Legumes of Malesia a Check-List: 1-295. Royal Botanic Gardens, Kew.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • Kumar, S. & Sane, P.V. (2003). Legumes of South Asia. A Checklist: 1-536. Royal Botanic Gardens, Kew.
    • Du Puy, D.J., Labat, N.-N., Rabevohitra, R., Villiers, J.-F., Bosser, J. & Moat, J. (2002). The Leguminosae of Madagascar: 1-737. Royal Botanic Gardens, Kew.
    • Mackinder, B., Pasquet, R., Polhill, R. & Verdcourt, B. (2001). Flora Zambesiaca 3(5): 1-261. Royal Botanic Gardens, Kew.
    • Stevens, W.D., Ulloa U., C., Pool, A. & Montiel, O.M. (2001). Flora de Nicaragua Monographs in Systematic Botany from the Missouri Botanical Garden 85: i-xlii, 1-2666. Missouri Botanical Garden.
    • Balick, M.J., Nee, M.H. & Atha, D.E. (2000). Checklist of the Vascular Plants of Belize with Common Names an Uses: 1-246. New Yourk Botanic Garden Press, New York.
    • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
    • Jørgensen, P.M. & León-Yánes, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador: 1-1181. Missouri Botanical Garden Press, St. Louis.
    • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
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    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

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    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    International Legume Database and Information Service
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    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Useful Plants of Boyacá Project
    ColPlantA database
    http://creativecommons.org/licenses/by-nc-sa/3.0
    http://creativecommons.org/licenses/by/3.0/