1. Family: Stilbaceae Kunth
    1. Genus: Nuxia Comm. ex Lam.
      1. Nuxia congesta R.Br. ex Fresen.

        This species is accepted, and its native range is Tropical & S. Africa, SW. Arabian Peninsula.

    [FWTA]

    Loganiaceae, C. F. A. Onochie & A. J. M. Leeuwenberg. Flora of West Tropical Africa 2. 1963

    Habit
    Tree up to 80 ft. high, sometimes with a grey fluted stem
    Flowers
    Flowers creamy-white and scented
    Ecology
    In montane forest.
    [FZ]

    Loganiaceae, A. J. M. Leeuwenberg. Flora Zambesiaca 7:1. 1983

    Habit
    Evergreen or sometimes in Ethiopia deciduous tree or shrub, 2–25 m. high, often gnarled.
    Trunk
    Trunk in larger trees fluted and irregularly ridged, 5–100(200) cm. in diam.; bark pale grey–brown, fissured.
    Branches
    Branchlets glabrous to densely pubescent, not lenticellate, with 6(8) raised lines and often sulcate when dry.
    Leaves
    Leaves ternate (occasionally 4–nate), petiolate; petiole glabrous to densely pubescent, 3–20 mm. long; lamina coriaceous or subcoriaceous, very variable in shape and size, elliptic, narrowly elliptic, obovate, nearly rhomboid, or suborbicular, 1·3–3(5) times as long as wide, (1)2–15 x (0·3)1·2–7·5 cm., acuminate to emarginate at the apex, cuneate or less often rounded at the base or decurrent into the petiole, entire or sometimes more or less distinctly serrate–dentate, or crenate, glabrous to subtomentose with stellate hairs on both sides. Leaves on sucker shoots (mostly in the shade) darker green, herbaceous, serrate, and more hairy in tropical countries, but thicker in southern Africa.
    Inflorescences
    Inflorescence variable in shape and size, seemingly umbellate to paniculate, congested or rather lax, 3–15 x 3–15 cm., 3–7 times branched. Peduncle and branches hairy like the branchlets.
    Pedicel
    Pedicels short or obsolete.
    Flowers
    Flowers solitary or three together, fragrant.
    Calyx
    Calyx green, often viscid, 3–8 x 1·5–2·2 mm., minutely to manifestly pubescent with glandular and mostly also ordinary hairs outside, inside sericeous; lobes variable in size, mostly about as long as wide or wider, 0·5–2 x 1–1·5 mm., acute or less often obtuse, entire, valvate, often two coherent or partially united.
    Corolla
    Corolla white, with recurved pubescence on the lobes which may be reduced to scattered hairs or is rarely absent and with minute glandular hairs at the base of the lobes and the apex of the tube (the latter sometimes on the middle of the tube or on nearly the complete tube too), inside with curved hairs in the throat and sometimes also at the apices of the lobes and with minute glandular hairs in the tube except for its glabrous base (hairs in throat or hairs in tube are occasionally absent, although never both simultaneously); tube slightly shorter or sometimes — when calyx very short only — longer than the calyx; lobes oblong, 2–3 x as long as wide, 2–5 x 1–2 mm., obtuse or subacute at the apex.
    Stamens
    Stamens: anthers glabrous, 1·2–2 mm. long.
    Pistil
    Pistil (4·5–)7·5–13 mm. long; ovary ovoid or ellipsoid, slightly laterally compressed, 1·5–2·5 x 1·2–2 x 1–1·5 mm., appressedly hirto–pubescent, glabrous at the base.
    Fruits
    Capsule 0·5–1·5 mm. longer than the calyx, appressedly hirto–pubescent.

    Images

    Distribution

    Native to:

    Angola, Botswana, Burundi, Cameroon, Cape Provinces, Eritrea, Ethiopia, Free State, Ghana, Guinea, Gulf of Guinea Is., Ivory Coast, Kenya, KwaZulu-Natal, Liberia, Malawi, Mozambique, Nigeria, Northern Provinces, Rwanda, Saudi Arabia, Sierra Leone, Somalia, Sudan, Swaziland, Tanzania, Uganda, Zambia, Zaïre, Zimbabwe

    Nuxia congesta R.Br. ex Fresen. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Nov 1, 2008 Onana, J.-M. [3634], Cameroon K000580276
    Dec 1, 2003 Tadjouteu, F. [426], Cameroon K000746063
    Jun 1, 2003 Cheek, M. [8972], Cameroon K000008624
    Jun 1, 2003 Etuge, M. [4382], Cameroon K000008623
    May 1, 2001 Cheek, M. [7345], Cameroon K000008625
    Dec 1, 2000 Etuge, M. [3460], Cameroon K000338856
    Jan 1, 1999 Buzgo, M. [632], Cameroon K000338855
    Jan 1, 1987 Manning, S.D. [2186], Cameroon K000028421
    Jan 1, 1977 Mott, P.J. [192], Botswana K000511152
    Jan 1, 1974 Zeyher [1327], South Africa K000426713 lectotype
    Jan 1, 1974 Chipp, T.F. [95], Sudan K000426758 holotype
    Jan 1, 1971 Scott Elliot, G.F. [4824], Sierra Leone K000426761 holotype
    Jan 1, 1971 Mann, G. [1206], Cameroon K000028434 isotype
    Jan 1, 1971 Mann, G. [1206], Cameroon K000028433 lectotype
    Jan 1, 1971 Mogg, A.O.D. [15186], South Africa K000426720 holotype
    Jan 1, 1971 Mogg, A.O.D. [15136], South Africa K000426721 holotype
    Jan 1, 1971 Schimper [1159], Ethiopia K000426759 isotype
    Jan 1, 1971 Schimper [1159], Ethiopia K000426760 isotype
    Jan 1, 1971 Welwitsch [5670], Angola K000426746 isotype
    Jan 1, 1971 K000426745 holotype
    Jan 1, 1971 Zeyher [1326], South Africa K000426718 isotype
    Jan 1, 1971 Zeyher [1326], South Africa K000426717 isotype
    Jan 1, 1971 Wood, J.M. [1862], South Africa K000426709 holotype
    Jan 1, 1971 Wood, J.M. [1862], South Africa K000426710 holotype
    Jan 1, 1971 Fries, R.E. [2035], Kenya K000426750 isotype
    Jan 1, 1971 Fries, R.E. [777], Kenya K000426751 isotype
    Jan 1, 1971 Fries, R.E. [759], Kenya K000426752
    Jan 1, 1971 Volkens, G. [1499], Tanzania K000426749 isotype
    Jan 1, 1971 Rogers, F.A. [19220], South Africa K000426711 holotype
    Jan 1, 1971 Rogers, F.A. [19220], South Africa K000426712 isotype
    Jan 1, 1971 Rogers, F.A. [23028], South Africa K000426715 isotype
    Jan 1, 1971 Rogers, F.A. [23028], South Africa K000426716 holotype
    Jan 1, 1971 Rogers, F.A. [6213], Botswana K000426747 isotype
    Jan 1, 1971 Rogers, F.A. [6213], Botswana K000426748 isotype
    Jan 1, 1971 Rogers, F.A. [6213], Botswana K000511155 Unknown type material
    Jan 1, 1971 Rudatis, H. [1399], South Africa K000426708 holotype
    Jan 1, 1971 McConnell, R.H. [68915], Botswana K000511151
    Jan 1, 1971 Hillary [543], Botswana K000511154
    May 10, 1970 Ash [395], Ethiopia 74525.000
    Jan 1, 1967 Adams, C.D. [GC 11217], Cameroon K000028436
    Jan 1, 1962 Breteler, F.J. [2573], Cameroon K000028415
    Jan 1, 1962 Breteler, F.J. [MC 140], Cameroon K000028418
    Jan 1, 1962 Breteler, F.J. [MC 140], Cameroon K000028420
    Jan 1, 1962 Lobe Babute, D. [Cam 15 /37], Cameroon K000028437
    May 1, 1959 Lightbody, J.S. [FHI 26281], Cameroon K000028425
    May 1, 1959 Lightbody, J.S. [FHI 26281], Cameroon K000028426
    Jan 1, 1959 Keay, R.W.J. [28630], Cameroon K000028422
    Jan 1, 1959 Maitland, T.D. [996], Cameroon K000028429
    Jan 1, 1959 Maitland, T.D. [458], Cameroon K000028430
    Jan 1, 1959 Maitland, T.D. [1194], Cameroon K000028431
    Jan 1, 1959 Mann, G. [2184], Cameroon K000028432
    Jan 1, 1959 Brenan, J.P.M. [9571], Cameroon K000028423
    Jan 1, 1959 Brenan, J.P.M. [9571], Cameroon K000028424
    Jan 1, 1959 Steele, M. [64A], Cameroon K000028428
    Drummond, R.B. [2626], Tanzania 16667.000
    Ash [738], Ethiopia 37177.000
    Hepper, F.N. [2203], Cameroon K000028427
    Gossweiler, J. [9307], Angola K001008574
    Newbould, J. [2326], Tanzania 16363.000
    Boughey, A.S. [12576], Cameroon K000028435
    Ash, J.W. [286], Ethiopia 7549.000
    Welwitsch [5678], Angola K001008575
    Welwitsch [5624], Angola K001008576
    Welwitsch [6689], Angola K001008577
    Rand [712], South Africa K000426719 isotype
    Breteler, F.J. [272], Cameroon K000028416
    Breteler, F.J. [166], Cameroon K000028419
    Hansen, O.J. [3206], Botswana K000511153
    Patel, H. [7364], Mozambique K000613626
    Timberlake, J. [5278], Mozambique K000614007
    Timberlake, J. [5105], Mozambique K000613485
    de Menezes, A. [1124], Angola K001008573
    Brunt, M.A. [102], Cameroon K000028417
    s.coll. [9206], Bahamas K001008570
    s.coll. [253], Angola K001008571
    s.coll. [257], Angola K001008572
    s.coll. [260], Angola K001008578

    First published in Flora 21: 606 (1838)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Velayos, M., Barberá, P., Cabezas, F.J., de la Estrella, M., Fero, M. & Aedo, C. (2014). Checklist of the vascular plants of Annobón (Equatorial Guinea) Phytotaxa 171: 1-78.
    • Jongkind, C. (2014). Fauna & Flora of Liberia, flowering plant species list www.liberianfaunaflora.org.
    • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Leeuwenberg, A.J.M. (1975). The Loganiaceae of Africa XIV. A revision of Nuxia Lam. Mededelingen Landbouwhogeschool te Wageningen 75-8: 1-80.

    Literature

    Flora of West Tropical Africa
    • Bruce & Lewis in F.T.E.A Loganiac. 44, fig. 8, 7-8 (1960).
    • F.T.A. 4, 1: 512 (1903)
    • in Flora 21: 606 (1838)
    Kew Backbone Distributions
    • Jongkind, C. (2014). Fauna & Flora of Liberia, flowering plant species list www.liberianfaunaflora.org.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Leeuwenberg, A.J.M. (1975). The Loganiaceae of Africa XIV. A revision of Nuxia Lam. Mededelingen Landbouwhogeschool te Wageningen 75-8: 1-80.

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0