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This species is accepted, and its native range is Eritrea to Zimbabwe, Arabian Peninsula.
A specimen from Kew's Herbarium

[FZ]

Rhamnaceae, R. B. Drummond. Flora Zambesiaca 2:2. 1966

Morphology General Habit
Shrub or tree to 7 m. tall; branches puberulous or glabrous, frequently spine-tipped; bark black.
Morphology Leaves
Leaves crowded at the end of short shoots; lamina 1·2–2·5 (3·5) × 0·7–1·5 cm., narrowly elliptic, elliptic, narrowly obovate or obovate, rounded and often emarginate at the apex, margin entire or obscurely denticulate, cuneate at the base, shining above, dull beneath, secondary nerves in 4–5 pairs, sparingly puberulous above; petiole 1–3 mm. long; stipules minute, ciliate.
Morphology Reproductive morphology Inflorescences
Inflorescences of few-flowered axillary fascicles.
Morphology Reproductive morphology Flowers
Flowers 4-merous, with pedicels 1–3 mm. long lengthening to 8–10 mm. long in fruit.
Morphology Reproductive morphology Flowers Receptacle
Receptacle 4-angled.
Morphology Reproductive morphology Flowers Calyx
Sepals 1·5 mm. long, deltate.
Morphology Reproductive morphology Flowers Corolla
Petals 1 mm. long, shorter than the stamens, hyaline.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens with filaments 1 mm. long.
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary 3-locular; 1 ovule per loculus; style short, 3-fid.
Morphology Reproductive morphology Fruits
Fruit 5 mm. in diam., somewhat fleshy.
Morphology Reproductive morphology Seeds
Seeds 3.

[FTEA]

Rhamnaceae, Marshall C. Johnston (University of Texas Herbarium). Flora of Tropical East Africa. 1972

Morphology General Habit
Deciduous shrubs or small trees, 0.5–7 m. tall, sometimes scandent.
Morphology General Bark
Bark smooth, grey; slash cream; year-old branches dark brown-grey, slender, glabrate; youngest twigs densely to less commonly sparsely hispidulous with grey hairs 0.1–0.2 mm. long; short shoots (many-noded twigs) often produced late in the growing season; elongated twigs occasionally thorn-tipped.
Morphology Leaves
Leaf-blades obovate or narrowly obovate or broadly oblanceolate, rarely elliptic, (1–)2–3(–4.4) cm. long, (5–)6–15(–20) mm. wide, cuneate, blunt to acute and mucronate, dark olive green or brown-green (at least when dry), glabrous or hispidulous on the major nerves beneath and occasionally above or rarely hispidulous on both surfaces with grey hairs 0.1–0.2 mm. long, obscurely glandular-serrulate or occasionally with the teeth reduced to vanishingly small size, with 5 to 8 pairs of obscure secondary nerves; petioles (1–)2–5(–8) mm. long, usually at first hispidulous and later glabrescent.
Morphology Leaves Stipules
Stipules filiform to subulate, 1–3 mm. long, caducous to subpersistent.
Morphology Reproductive morphology Flowers
Flowers borne singly at the nodes, often appearing at the nodes of the short-shoots of the previous season and thus in somewhat fascicle-like aggregations, greenish or sometimes said to be cream colour or orange, fragrant; pedicels usually glabrous, 1–2 (–4) mm. long in flower, in fruit ± 2(–4.5) mm. long.
Morphology Reproductive morphology Flowers Hypanthium
Cup glabrous or rarely hispidulous.
Morphology Reproductive morphology Flowers Calyx
Sepals 4, deltoid, 1–1.7 mm. long, glabrous or rarely hispidulous on back.
Morphology Reproductive morphology Flowers Corolla
Petals 4, linear to oblanceolate, ± 0.5–0.8 mm. long, not clasping, caducous.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 4, said to be brownish when fresh.
Morphology Reproductive morphology Fruits
Fruit nearly globose, ± 5–5.5 mm. long, reddish or scarlet when ripe, with 3 stones or by abortion fewer, each stone on drying eventually capable of colubrinoid dehiscence, revealing an orange or brick red inner lining.
Morphology Reproductive morphology Seeds
Seed ± 4–5 mm. long, pale brownish, smooth, with a ventral groove.
Habitat
Forest margins and upland evergreen bushland; 1000–3000 m. often at high altitudes
Distribution
K1 K3 K4 K5 K6 T1 T2 T3 T7 U2 U4

[FSOM]

M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS

Morphology General Habit
Evergreen shrub or small tree; branches sometimes spine-tipped, sparsely pubescent when young
Morphology Leaves
Leaves alternate, leathery, glabrous to pubescent on petiole and on major nerves; petiole 1–11 mm long; blade oblanceolate to elliptic or narrowly obovate, 1–11 mm long; blade oblanceolate to elliptic or narrowly obovate, 1–5.5 x 0.4–3 cm, glandular-serrulate to entire, acute to rounded at the apex, attenuate to cuneate at the base
Morphology Reproductive morphology Flowers
Flowers yellowish-green, solitary or in 2–4-flowered clusters, 4-merous; pedicels 1–5 mm long
Morphology Reproductive morphology Flowers Calyx
Sepals 1.5–2 mm long
Morphology Reproductive morphology Flowers Corolla
Petals 0.5–1 mm long, soon falling
Morphology Reproductive morphology Flowers Gynoecium Style
Style c. 1 mm long
Morphology Reproductive morphology Fruits
Drupe subglobose, 4–7 mm in diam., first red then blackish purple.
Distribution
N1–3 Eritrea, Ethiopia and southwards to Zimbabwe.
Ecology
Altitude range 1100–2175 m.
Vernacular
Alendondoh, jajale, madowya (Somali).

Native to:

Burundi, Djibouti, Eritrea, Ethiopia, Kenya, Oman, Rwanda, Somalia, Tanzania, Uganda, Yemen, Zaïre, Zimbabwe

Rhamnus staddo A.Rich. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Jan 1, 1961 Dehn, G. [R40], Zimbabwe K000036310 isotype

First published in Tent. Fl. Abyss. 1: 138 (1847)

Accepted by

  • Exell, A.W., Fernandes, A. & Wild, H. (eds.) (1966). Flora Zambesiaca 2(2): 353-653. Royal Botanic Gardens, Kew.
  • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
  • Ghazanfar, S.A. (1992). An Annotated Catalogue of the Vascular Plants of Oman and their Vernacular names Scripta Botanica Belgica 2: 1-153.
  • Ghazanfar, S.A. (2007). Flora of Oman 2: 1-220. National Botanic Garden of Belgium, Meise.
  • Hedberg, I. & Edwards, S. (eds.) (1989 publ. 1990). Flora of Ethiopia and Eritrea 3: 1-659. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Johnston, M.C. (1972). Flora of Tropical East Africa, Rhamnaceae: 1-40.
  • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
  • Mosti, S., Raffaelli, M. & Tardelli, M. (2012). Contributions to the flora of central-southern Dhofar (Sultanate of Oman) Webbia; Raccolta de Scritti Botanici 67: 65-91.
  • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.
  • Thulin, M. (ed.) in Thulin, M. (ed.) (1999). Flora of Somalia 2: 1-303. The Royal Botanic Gardens, Kew.
  • Troupin, G. (ed.) (1983). Flora du Rwanda 2: 1-603. Musee Royal de l'Afrique Centrale.
  • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.

Literature

Kew Backbone Distributions

  • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
  • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
  • Hedberg, I. & Edwards, S. (eds.) (1989 publ. 1990). Flora of Ethiopia and Eritrea 3: 1-659. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Mosti, S., Raffaelli, M. & Tardelli, M. (2012). Contributions to the flora of central-southern Dhofar (Sultanate of Oman) Webbia; Raccolta de Scritti Botanici 67: 65-91.

Flora of Somalia

  • Flora Somalia, Vol 2, (1999) Author: by M. Thulin [updated by M. Thulin 2008]

Flora of Tropical East Africa

  • A. Rich., Tent. Fl. Abyss. 1: 138 (1847).
  • Dale & Greenway, Kenya Trees and Shrubs p. 391, fig. 77 (1961).
  • Evrard in Flore du Congo Belge et du Ruanda-Urundi, 9: 434 (1960).
  • Hemsl. in Flora of Tropical Africa 1: 382 (1868).
  • R.B. Drummond in Flora Zambesiaca 2: 428, t. 89/C (1966).

Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Somalia
Flora of Somalia
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0