1. Family: Rubiaceae Juss.
    1. Genus: Coffea L.
      1. Coffea arabica L.

        Coffee is the world's favourite drink, the most important commercial crop-plant, and the second most valuable international commodity after oil. Worldwide there are about 20 million coffee farming families; around 100 million people depend on coffee for their livelihoods. Its export value alone is immense (US $ 15.4 billion in 2009/10) and as such it plays a crucial role in the economies of several tropical countries.

    [KSP]

    Kew Species Profiles

    General Description

    Coffee is the world's favourite drink, the most important commercial crop-plant, and the second most valuable international commodity after oil. Worldwide there are about 20 million coffee farming families; around 100 million people depend on coffee for their livelihoods. Its export value alone is immense (US $ 15.4 billion in 2009/10) and as such it plays a crucial role in the economies of several tropical countries.

    Coffee is made from the roasted seeds of the genus Coffea, and is brought from plant to cup via a complex process. The glossy red fruits are picked, the fleshy outer part of the berry removed and then their pale-coloured seeds (beans) are sent to mills to remove the hard outer layer encasing the seeds. At this stage the beans are either exported, or they are blended with others from the region before exportation. The complicated process of roasting usually occurs in the country where the beverage is consumed.

    Species Profile
    Geography and distribution

    Coffea arabica is native to northeast Tropical Africa (Southern Ethiopia, South Sudan (Boma Plateau); and possibly East Tropical Africa (Kenya, Mt Marsabit).  It is sometimes naturalised in tropical areas. Arabica coffee grows at 950 m to 1,950 m above sea level.

    In total, there are 125 coffee species, which occur naturally in Africa, the Indian Ocean Islands, (Madagascar, Comoros, and the Mascarenes), southern Asia, south east Asia and Australia.  Arabica and Robusta coffee are the main beverage species, with a small percentage of Liberica coffee (Coffea liberica ) grown for commercial purposes. Other species in East Africa and Madagascar are sometimes used locally to make coffee on a very small scale.

    Description

    Overview: A small tree, 2 to 8 m tall, with characteristic horizontal branching, although in plantations it is usually pruned to take the form of a small shrub. Its leaves are evergreen and usually shiny.

    Flowers: The flowers are hermaphroditic and sweet-scented, the corolla white, tubular, normally with 5 lobes.

    Fruits: The fruits are usually red but sometimes yellow or purple at maturity. The outer layer is soft, edible and sweet-tasting, containing two or sometimes one 'seed' — actually a seed encased in a hard, almost crispy outer layer which together forms a structure known as a pyrene; this outer layer is removed by milling.

    Seeds: The seed itself is pale fawn or brown (dark brown only after roasting) and has a characteristic groove on its inner surface, which curls round inside the seed.

    Arabica coffee is a hybrid species, formed by the hybridization of Coffea eugenioides and Coffea canephora . It is one of the only species in the genus Coffea that is self-fertile (autogamous), a single plant being able to produce fertile seed from its own pollen.

    Threats and conservation

    Based on its extent of occurrence alone, Arabica coffee falls within the Vulnerable extinction risk category. However, a rating of Endangered is more likely, owing to ongoing deforestation in Ethiopia and elsewhere, and the threats posed by climate change. Arabica is a climate-sensitive species, being found only in humid, montane forests at 950-1950 m. The montane forests of south-western Ethiopia are by far the most important area in terms of natural distribution and genetic diversity.

    Cultivated Arabica is under threat because of a low-level of genetic diversity within the crop cultivars, leaving plantations vulnerable to pests, diseases and climate change. These problems are compounded for both wild populations and crops because coffee seeds cannot yet be stored successfully in conventional seed banks (in low temperature, low moisture environments). In vitro germplasm collections, cryopreservation and living collections are possible alternatives but they are expensive and time-consuming to maintain.

    Kew produces conservation assessments for coffee species as part of its important coffee research programme.

    Coffee plantations can be managed to promote biodiversity, with many plantations now being certified by fair-trade and forest alliance organisations. In particular, shade coffee can be planted with forest trees, to promote wildlife and particularly bird species.

    Uses

    Coffee cultivation may have started in the sixth century in Yemen, its use being for spiritual purposes.

    Coffee became a popular drink in Europe from the seventeenth century onwards, being imported from plantations established first by the Dutch in Ceylon (now Sri Lanka), Java, and later from plantations in Brazil and the West Indies established in the eighteenth century. Much of the world's Arabica coffee is produced in Latin America.

    The stimulating effects of coffee are largely due to the alkaloid caffeine contained in the seeds. As well as a beverage, coffee is used for flavouring foods and confectionery. The beans are also a commercial source of caffeine, a by-product of making de-caffeinated coffee. Caffeine is added to soft drinks and medicines as a stimulant and diuretic. Roasted and ground coffee is a constituent of traditional medicines in South-East Asia to alleviate stomach ache and diarrhoea, to increase blood pressure, and as a diuretic. In some countries coffee leaves are used to make a hot drink, like tea.

    Coffee wood, from the main trunk, is used locally in construction. David Livingstone, the nineteenth century explorer and missionary, reported seeing coffee trees being used to make huts in his travels in southern Africa. The timber is straight, dense, strong and partially resistant to termites. The wood is also used for furniture and as fuel wood.

    Arabica and Robusta coffee

    Originally from the high altitude, humid forests of Ethiopia and South Sudan, where it still grows wild, Arabica coffee ( Coffea arabica ) is considered to produce the finest coffee beans. Most instant coffee is made from a mixture of Arabica and Robusta ( Coffea canephora ), the latter having a less delicate flavour and aroma and more caffeine. Generally, Arabica is grown in upland plantations and Robusta in lowland plantations.

    Millennium Seed Bank: Seed storage

    Kew's Millennium Seed Bank Partnership aims to save plant life worldwide, focusing on plants under threat and those of most use in the future. Seeds are dried, packaged and stored at a sub-zero temperature in our seed bank vault.

    You can find out more about storing and germinating the seeds of Coffee arabica in Kew's Difficult Seeds Project

    Distribution
    Ethiopia, Kenya
    Ecology
    Humid, evergreen forest.
    Conservation
    Vulnerable (provisional assessment).
    Hazards

    Although recent research shows that there are many positive health benefits from consumption in moderation, much research is being undertaken to investigate the numerous compounds found in coffee and how these affect quality and human health.

    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Bush or tree 2–7 m. tall; young branches glabrous, covered with light brown shiny bark.
    Leaves
    Leaf-blades elliptic to broadly elliptic or oblong-elliptic (or lanceolate or round in some cultivated forms), 7–18 cm. long, 3–7.5 cm. wide, distinctly acuminate at apex, acute to obtuse at base, thinly coriaceous to coriaceous, with 7–10 pairs of main lateral nerves, shiny above, margin sometimes undulate; domatia rather inconspicuous, glabrous, occasionally absent from some leaves; stipules triangular, 4–8 mm. long, acute, usually exceeded by a mucro.
    Flowers
    Flowers (4–)5(–6)-merous, 2–20 per axil, borne in 1–3(–15) fascicles; peduncle 0.5–2(–3) mm. long in flower (2–4 mm. long in fruit); pedicels 1–2(–3) mm. long (calyces usually well clear of surrounding bracteole); bracteoles frequently with subfoliaceous lobes up to 6 mm. long.
    Calyx
    Calyx-tube 1–2 mm. long; limb reduced to a rim or sometimes irregularly toothed; teeth up to 1(–2) mm. long.
    Corolla
    Corolla-tube (0.5–)0.9–1.1 cm. long, 2–3 mm. wide at throat; lobes oblong, 0.9–1.6 cm. long, (2–)4–6 mm. wide, rounded.
    Fruits
    Fruit red (yellow or purple in cultivated forms), oblong-ellipsoid or sometimes ± subglobose, 1–2 cm. long, 0.9–1.1 cm. wide; pedicel lengthening to 4–8 mm.
    Seeds
    Seeds fawn or greenish fawn, 9–1.2(–1.5) cm. long, 6–7 mm. wide.
    Habitat
    Forest; 1370–1525 m.
    Distribution
    K1 widely cultivated throughout the tropics

    Images

    Distribution

    Native to:

    Ethiopia, Kenya, Sudan

    Introduced into:

    Andaman Is., Ascension, Bangladesh, Belize, Bermuda, Bismarck Archipelago, Bolivia, Borneo, Brazil North, Brazil South, Brazil Southeast, Brazil West-Central, Central American Pac, China South-Central, China Southeast, Colombia, Congo, Cook Is., Costa Rica, Cuba, Dominican Republic, Easter Is., Ecuador, El Salvador, Fiji, Gabon, Galápagos, Guatemala, Guinea-Bissau, Gulf of Guinea Is., Hainan, Haiti, Honduras, Jamaica, Leeward Is., Lesser Sunda Is., Malawi, Marquesas, Mexico Central, Mexico Southeast, Mexico Southwest, Myanmar, New Caledonia, Nicaragua, Norfolk Is., Ogasawara-shoto, Panamá, Peru, Puerto Rico, Rwanda, Society Is., St.Helena, Taiwan, Trinidad-Tobago, Tubuai Is., Venezuela, Windward Is.

    Common Names

    English
    Arabica coffee

    Coffea arabica L. appears in other Kew resources:

    Date Identified Reference Herbarium Specimen Type Status
    Oct 1, 1969 Mooney, H.F. [5733], Ethiopia K000912159
    32258.000
    Small, D. [248], Sierra Leone 36222.000
    s.coll. [Cat. no. 6243], India K001123398
    s.coll. [Cat. no. 6243] K001123399
    s.coll. [Cat. no. 6243] K001123400
    s.coll. [Cat. no. 6243] K001123401
    s.coll. [Cat. no. 6243] K001123402
    s.coll. [Cat. no. 6243], India K001123403

    First published in Sp. Pl.: 172 (1753)

    Accepted by

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    • Baksh-Comeau, Y., Maharaj, S.S., Adams, C.D., Harris, S.A., Filer, D.L. & Hawthorne, W.D. (2016). An annotated checklist of the vascular plants of Trinidad and Tobago with analysis of vegetation types and botanical 'hotspots' Phytotaxa 250: 1-431.
    • Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Hammel, B.E., Grayum, M.H., Herrera, C. & Zamora, N. (eds.) (2014). Manual de Plantas de Costa Rica 7: 1-840. Missouri Botanical Garden Press, St. Louis.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • López Patiño, E.J., Szeszko, D.R., Rascala Pérez, J. & Beltrán Retis, A.S. (2012). The flora of the Tenacingo-Malinalco-Zumpahuacán protected natural area, state of Mexico, Mexico Harvard Papers in Botany 17: 65-167.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • Lambdon, P. (2012). Flowering plants & ferns of St Helena: 1-624. Pisces publications for St Helena nature conservation group.
    • Figueiredo, E., Paiva, J., Stévart, T., Oliveira, F. & Smith, G.F. (2011). Annotated catalogue of the flowering plants of São Tomé and Príncipe Bothalia 41: 41-82.
    • Wu, Z., Raven, P.H. & Hong, D. (eds.) (2011). Flora of China 19: 1-884. Missouri Botanical Garden Press, St. Louis.
    • Delprete, P.G. (2010). Rubiaceae Flora dos estados de Goiás e Tocantins 40: 1-1610. Universidade Federal de Goiás.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Davis, A.P., Govaerts, R., Bridson, D.M. & Stoffelen, P. (2006). An annotated taxonomic conspectus of the genus Coffea (Rubiaceae) Botanical Journal of the Linnean Society 152: 465-512.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Strugnell, A.M. (2006). A checklist of the Spermatophytes of Mt. Mulanje, Malawi Scripta Botanica Belgica 34: 1-199.
    • Borhidi, A. (2006). Rubiáceas de México: 1-512. Akadémiai Kiadó, Budapest.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
    • Takeuchi, W. (2005). Floristic notes from a holocene successional environment in Papuasia Harvard Papers in Botany 10: 95-116.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Delprete, P.G., Smith, L.B. & Klein, R.M. (2004). Flora Ilustrada Catarinense 1: 1-344. Herbário "Barbosa Rodrigues", Atají, Brasil.
    • Beaman, J.H. & Anderson, C. (2004). The Plants of Mount Kinabalu 5: 1-609. Natural history publications (Borneo), Kota Kinabalu, Malaysia.
    • Fairhurst, W. (2004). Flowering Plants of Ascension island: 1-300. Higham Press, Shirland, Alfreton, England.
    • Mendoza, H., Ramirez P., B.R. & Jimenez, L.C. (2004). Rubiaceae de Colombia: guia ilustrada de generos: 1-351. Instituto de Investigacion de Recursos Biologicos Alexander von Humboldt, Bogota, Colombia.
    • Berry, P.E., Yatskievych, K. & Holst, B.K. (eds.) in Steyermark, J., Berry, P.E., Holts, B.K. (eds). (2004). Flora of the Venezuelan Guayana 8: 1-874. Missouri Botanical Garden, St. Louis.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
    • Welsh, S.L. (1998). Flora Societensis: 1-420. E.P.S. Inc. Utah.
    • Orchard, A.E. (ed.) (1994). Oceanic Islands 1 Flora of Australia 49: 1-681. Australian Government Publishing Service, Canberra.
    • MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
    • Smith, A.C. (1988). Flora Vitiensis Nova. A new flora for Fiji (Spermatophytes only) 4: 1-377. Pacific Tropical Botanical Garden, Lawai.

    Literature

    Kew Species Profiles
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    Kew Backbone Distributions
    • Jaramillo Díaz, P. & Guézou, A. (2017). CDF Checklist of Galapagos Vascular Plants - FCD Lista de especies de Plantas Vasculares de Galápagos http://www.darwinfoundation.org/datazone/checklists/vascular-plants/.
    • Baksh-Comeau, Y., Maharaj, S.S., Adams, C.D., Harris, S.A., Filer, D.L. & Hawthorne, W.D. (2016). An annotated checklist of the vascular plants of Trinidad and Tobago with analysis of vegetation types and botanical 'hotspots' Phytotaxa 250: 1-431.
    • Sykes, W.R. (2016). Flora of the Cook Islands: 1-973. National Tropical Botanical Garden, Hawaii.
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    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
    • Lambdon, P. (2012). Flowering plants & ferns of St Helena: 1-624. Pisces publications for St Helena nature conservation group.
    • Figueiredo, E., Paiva, J., Stévart, T., Oliveira, F. & Smith, G.F. (2011). Annotated catalogue of the flowering plants of São Tomé and Príncipe Bothalia 41: 41-82.
    • Wu, Z., Raven, P.H. & Hong, D. (eds.) (2011). Flora of China 19: 1-884. Missouri Botanical Garden Press, St. Louis.
    • Delprete, P.G. (2010). Rubiaceae Flora dos estados de Goiás e Tocantins 40: 1-1610. Universidade Federal de Goiás.
    • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Delprete, P.G. & Cortés-B., R. (2006 publ. 2007). A synopsis of the Rubiaceae of the states of Mato Grosso and Mato Grosso do Sul, Brazil, with a key to genera, and a preliminary species list Revista de Biologia Neotropical 3: 13-96.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Strugnell, A.M. (2006). A checklist of the Spermatophytes of Mt. Mulanje, Malawi Scripta Botanica Belgica 34: 1-199.
    • Catarino, L., Sampaio Martins, E., Pinto-Basto, M.F. & Diniz, M.A. (2006). Plantas Vasculares e Briófitos da Guiné-Bissau: 1-298. Instituto de investigação científica tropical, Instituto Português de apoio ao desenvolvimento.
    • Takeuchi, W. (2005). Floristic notes from a holocene successional environment in Papuasia Harvard Papers in Botany 10: 95-116.
    • Tanaka, N., Koyama, T. & Murata, J. (2005). The flowering plants of Mt. Popa, central Myanmar - Results of Myanmar-Japanese joint expeditions, 2000-2004 Makinoa 5: 1-102.
    • Delprete, P.G., Smith, L.B. & Klein, R.M. (2004). Flora Ilustrada Catarinense 1: 1-344. Herbário "Barbosa Rodrigues", Atají, Brasil.
    • Beaman, J.H. & Anderson, C. (2004). The Plants of Mount Kinabalu 5: 1-609. Natural history publications (Borneo), Kota Kinabalu, Malaysia.
    • Fairhurst, W. (2004). Flowering Plants of Ascension island: 1-300. Higham Press, Shirland, Alfreton, England.
    • Berry, P.E., Yatskievych, K. & Holst, B.K. (eds.) in Steyermark, J., Berry, P.E., Holts, B.K. (eds). (2004). Flora of the Venezuelan Guayana 8: 1-874. Missouri Botanical Garden, St. Louis.
    • Welsh, S.L. (1998). Flora Societensis: 1-420. E.P.S. Inc. Utah.
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    Sources

    Art and Illustrations in Digifolia
    [A] Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    [B] http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    [C] © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2018. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    [D] © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Species Profiles
    Kew Species Profiles
    [E]
    [F] http://creativecommons.org/licenses/by-nc-sa/3.0