1. Family: Rubiaceae Juss.
    1. Genus: Kohautia Cham. & Schltdl.
      1. Kohautia aspera (B.Heyne ex Roth) Bremek.

        This species is accepted, and its native range is Cape Verde, Dry Tropical & S. Africa, Arabian Peninsula, Pakistan to India.

    [FZ]

    Rubiaceae, B. Verdcourt. Flora Zambesiaca 5:1. 1989

    Habit
    Erect unbranched to sparsely branched annual herbs 10–30 cm. tall; stems ± densely papillose often reddish at base and below nodes.
    Leaves
    Leaves (23)28–39(47) x (0.5)0.7–2.5(4) mm., linear, linear-lanceolate to narrowly elliptic-lanceolate, margins and midrib beneath papillose, otherwise surfaces scabrid to glabrous, apex acute, narrowed to the base; stipular sheath (0.7)1–1.6(2) mm. long, stipular lobes (0.5)0.7–1.2(2.5) mm. long.
    Inflorescences
    Inflorescences spike-like, many-flowered, mostly 2 subsessile flowers at a node, the flower in a fork of a dichasium distinctly pedicellate; peduncles 1–60 mm. long; pedicels (0)2–10(25) mm. long, all parts densely papillose like the stem.
    Calyx
    Calyx tube (0.6)1–1.4(1.5) x (0.5)0.7–1.5(1.7) mm. ovoid to elliptic, densely covered with rounded papillae; lobes (0.8)1–1.4(1.7) mm. long and (0.2)0.3–0.5(0.6) mm., narrowly triangular to linear-lanceolate, margins and midrib scabrid with stiff minute papillae.
    Corolla
    Corolla lobes above mostly white or blueish or pinkish white, bluish or mauve below, tube light green or greeny white, narrow part of tube outside with rounded minute papillae; tube slightly curved, altogether (2.1)2.3–3.1(3.4) mm. long, the dilated part ovoid-elliptic in shape, (0.8)1–1.3(1.5) x (0.4)0.5–0.8(0.9) mm., the narrow part (0.2)0.3–0.6(0.7) mm. wide; lobes (0.7)1–1.5(1.8) x (0.3)0.4–0.7(0.9) mm., ovate-triangular to ovate-oblong, not parted to the base, acute.
    Anthers
    Anthers ± oval, c. two thirds the length of the dilated tube; pollen 4–5-colporate.
    Pistil
    Style and stigma as long as the narrow part of the tube, stigma lobes 0.7–1(1.4) mm. long, tips touching the anthers.
    Fruits
    Capsules (2)2.2–3(3.4) x (2.8)3.4–4.3(4.7) mm., subglobose to globose, ± sparsely papillose.
    Seeds
    Seeds light brown, c. 0.4–0.5 mm. long, angular-subconic.
    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Annual sparsely to densely branched herb 13–40 cm. tall, the stems scabrid with white papillae.
    Leaves
    Leaves linear to narrowly elliptic, 1.5–3(–4.7) cm. long, 0.5–4 mm. wide, acute at the apex, narrowed to the base, glabrous above, papillate on midvein beneath and on the margins; stipule-sheath 0.7–1.5 mm. long, divided into 2 fimbriae 0.5–2 mm. long.
    Inflorescences
    Inflorescences lax, the flowers mostly in pairs well spaced on the rhachis, entirely monochasial or branched at the base, trichotomous or dichasial; peduncle 1–4(–8) cm. long; pedicels 0.1–1.3(–2.6) cm. long; all parts papillate like the stems.
    Calyx
    Calyx-tube ellipsoid, 1–1.2 mm. long, papillate; lobes linear-lanceolate, 0.7–2 mm. long, glabrous or minutely scabridulous.
    Corolla
    Corolla white, brownish, greenish, pale blue or purple; tube narrowly cylindrical, sometimes curved, 2.5–4.7 mm. long, the widened part 0.7–2 mm. long, 0.5–0.8 mm. wide, mostly sparsely papillate outside with downwardly directed hairs inside near the base of the widened part; lobes oblong, 1–1.2 mm. long, 0.5–0.9 mm. wide.
    Pistil
    Style ± 0.6 mm. long; stigma-lobes 1.2–1.4 mm. long.
    Fruits
    Capsule oblong-subglobose, 2.5–3(–4) mm. tall, 2.5–4 mm. wide, ± smooth or papillate, the dehisced beak slightly raised.
    Seeds
    Seeds pale brown, angular, 0.4–0.6 mm. long.
    Habitat
    Grassland, open bushland and thicket, also overgrazed wet ground and as a weed in arable fields, particularly on heavy black clay soils; 750–1950 m.
    Distribution
    South West Africa K1 K3 K4 K6 T1 T2 T3 T4 T5 T7 U1

    Images

    Distribution

    Native to:

    Botswana, Cape Verde, Chad, Djibouti, Eritrea, Ethiopia, Free State, India, Kenya, Mali, Mauritania, Namibia, Niger, Northern Provinces, Pakistan, Senegal, Somalia, Sudan, Tanzania, Uganda, Yemen, Zimbabwe

    Kohautia aspera (B.Heyne ex Roth) Bremek. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1950 Kotschy [46], Ethiopia K000172598 Unknown type material
    s.coll. [Cat. no. 865] Hedyotis elongata K001110087
    s.coll. [Cat. no. 865] Hedyotis elongata K001110088
    s.coll. [Cat. no. 865] Hedyotis elongata K001110089
    s.coll. [Cat. no. 865], India Hedyotis elongata K001110090
    s.coll. [Cat. no. 865], India Hedyotis elongata K001110091

    First published in Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Sect. 2, 48(2): 113 (1952)

    Accepted by

    • Balkrishna, A. (2018). Flora of Morni Hills (Research & Possibilities): 1-581. Divya Yoga Mandir Trust.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Lebrun, J.-P. (1998). Catalogue des plantes de la Mauritanie et du Sahara Occidental Boissiera 55: 1-322.
    • Lebrun, J.P. (1973). Énumération des plantes vasculaires du Sénégal: 1-209. Maisons Alfort: Institut d'élevage et de médecine vétérinaire des pays tropicaux.

    Literature

    Kew Backbone Distributions
    • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
    • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
    • Ehrendorfer, F., Schönbeck-Temesy, E., Puff, C. & Rechinger, W. (2005). Rubiaceae Flora Iranica 176: 1-287. Naturhistorisches Museums Wien.
    • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
    • Lebrun, J.P. (1973). Énumération des plantes vasculaires du Sénégal: 1-209. Maisons Alfort: Institut d'élevage et de médecine vétérinaire des pays tropicaux.
    Flora of Tropical East Africa
    • A.D.Q. Agnew, Upland Kenya Wild Flowers p. 398 (1974).
    • Lebrun in Mitt. Bot. Staats., München 10: 444, fig. 5 (1971).
    • Bremek. in Verh. K. Nederl. Akad. Wet., Afd. Natuurk., ser. 2, 48(2): 113 (1952).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0