1. Family: Apiaceae Lindl.
    1. Genus: Berula W.D.J.Koch
      1. Berula imbricata (Schinz) Spalik & S.R.Downie

        This species is accepted, and its native range is Cameroon to W. & S. Tanzania and S. Tropical Africa.

    [FZ]

    Umbelliferae, J. F. M. Cannon. Flora Zambesiaca 4. 1978

    Habit
    Glabrous herb 30–100 cm. tall.
    Roots
    Rootstock probably a creeping rhizome with tuberous nodules from which adventitious roots arise at the nodes.
    Stem
    Stem terete, hollow with a large central canal, finely grooved, unbranched below the level of the inflorescence.
    Leaves
    Leaves up to 35 cm. long, simply pinnate, petiole and rhachis terete, fistulose and septate; leaflets rigid, somewhat coriaceous, irregularly ovoid with a cartilaginous margin and strongly developed teeth. Leaf morphology very variable, probably in association with habitat conditions; in extreme cases the leaves may be reduced to little more than fistulose petioles. In young leaves the leaflets are condensed and imbricated.
    Inflorescences
    Inflorescence sparsely branched. Umbels compound with up to 25 rays; bracts up to 10 × 2 mm., linear-lanceolate, numerous, becoming subulate at the apex and with distinct membranous margins. Partial umbels with 10–25 flowers on thin, unequal pedicels; bracteoles 8–10, similar to the bracts.
    Corolla
    Petals greenish-white to yellow.
    Fruits
    Fruit (not fully mature) 2 × 1 mm., slightly compressed laterally, primary ribs well developed and conspicuous; Stylopodium conical, yellowish; styles 0·25 mm. long, brownish; vittae 1 in each interval and 2 in the commissural face.
    [FTEA]

    Umbelliferae, C.C. Townsend. Flora of Tropical East Africa. 1989

    Habit
    Glabrous perennial herb 22–100 cm. tall, with a thickened, dark, woody rhizome sending out numerous fibrous roodets and occasionally hypogeal stolons.
    Stem
    Stem fistular, strongly ridged and sulcate, with generally few, slender, long, ascending branches.
    Leaves
    First leaves generally of a single oblong to elliptic leaflet with 1–2 pairs of rudimentary lower leaflets, other lower leaves with an increasing number of leaflets; main lower leaves linear-oblong in oudine, up to 70 cm. long but generally less, simply pinnate with 16–32(–40) pairs of firm, roundish to oblong, flabellate or linear-falcate, ± 5–20 × 3–15 mm. leaflets; leaflets denticulate with broad-based teeth ± abruptly narrowed to a sharp, commonly whitish cartilaginous mucro, the lowest on the fistular rhachis often reduced and rudimentary; sheaths striate, narrowed above, membranous-margined, 0.8–2.5 cm.; petiole 0.6–5 cm.; upper leaves rapidly reduced, becoming very small, sessile, with 2–4 pairs of mucronate to subaristate segments.
    Inflorescences
    Umbels on 2–14 cm. peduncles; rays 5–10(–25), slender, 1–7 cm., glabrous; involucre conspicuous, of 4–10 bracts which are broadly pale-margined below, entire or usually trisect or pinnate above, 0.8–1.6 cm., finally deflexed; partial umbels ± 10–28-flowered, pedicels glabrous, ± 2–8 mm.; involucel of 5–8 broadly elliptic, acute to acuminate, broadly pale-margined, finally deflexed, 3–5 mm. bracteoles.
    Calyx
    Sepals very small, sometimes not discernible.
    Corolla
    Petals cream or white to greenish-yellow, 0.75–1 mm.
    Fruits
    Fruit broadly ovoid, ± 2.5 × 2 mm., glabrous, blackish or dark red, the mericarps sometimes gaping centrally at the commissure when ripe, ribs narrow but pale and prominent; stylopodia large, shortly conical, subequalled by the rather rigid, ± divergent styles.
    Figures
    Fig. 23.
    Habitat
    On peaty soil in bogs or swamps, especially along rivers, also in damp montane grassland; 960–2100 m.
    Distribution
    T7 T8

    Distribution

    Native to:

    Angola, Cameroon, Central African Repu, Malawi, Mozambique, Tanzania, Zambia, Zaïre, Zimbabwe

    Berula imbricata (Schinz) Spalik & S.R.Downie appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jun 1, 2000 Cheek, M. [9889], Cameroon Afrocarum imbricatum K000338819

    First published in Taxon 58: 745 (2009)

    Accepted by

    • Van Wyk, B.-E., Tilney, P.M. & Magee, A.R. (2013). African Apiaceae: a synopsis of the Apiaceae/Umbelliferae of Sub-Saharan Africa and Madagascar: 1- 317. Briza Academic Books, Pretoria.

    Literature

    Kew Backbone Distributions
    • Van Wyk, B.-E., Tilney, P.M. & Magee, A.R. (2013). African Apiaceae: a synopsis of the Apiaceae/Umbelliferae of Sub-Saharan Africa and Madagascar: 1- 317. Briza Academic Books, Pretoria.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0