1. Family: Rubiaceae Juss.
    1. Genus: Bullockia (Bridson) Razafim., Lantz & B.Bremer
      1. Bullockia mombazensis (Baill.) Razafim., Lantz & B.Bremer

        This species is accepted, and its native range is S. Somalia to Mozambique.

    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Compact, or less often straggling, shrub or small tree 2–7 m. tall (or rarely a ? liane); bark pale grey, glabrous or occasionally pubescent.
    Leaves
    Leaf-blades pale greyish to bronze above when dry, broadly elliptic to round or broadly oblong-elliptic, 2.2–13 cm. long, 1–7.5 cm. wide, obtuse to rounded at apex, obtuse to rounded or sometimes truncate at base, often unequal, glabrous or sometimes pubescent beneath; lateral nerves apparent, in 4 main pairs; tertiary nerves always obscure; domatia absent or inconspicuous; petiole 0.2–1 cm. long; stipules light brown when dry, ovate or sometimes narrowly ovate, 0.4–1.6 cm. long, 0.2–1 cm. wide, acuminate.
    Flowers
    Flowers 5–6-merous. Functionally ♂ flowers borne in 3–20-flowered, sessile or rarely pedunculate (up to 2 mm.) umbels; pedicels 3–7 mm. long, glabrous or sometimes pubescent; calyx-tube reduced; limb lobed almost to base; lobes somewhat unequal, linear-triangular, occasionally slightly spathulate, 0.75–2 mm. long, often with a few hairs at apex; corolla whitish or yellowish green, readily caducous; tube ± 1.5 mm. long, with a ring of deflexed hairs at throat inside; lobes triangular-ovate, 1.25–1.5 mm. long, 1–1.5 mm. wide at base, acute, often with a few hairs at apex outside, erect; anthers exserted but almost sessile, 0.75 mm. long; style shortly exceeding corolla-tube; stigmatic knob 0.25 mm. long and wide. Functionally ♀ flowers solitary; pedicels 0.4–1 cm. long; calyx-tube 1.75–2.5 mm. long; lobes more strongly spathulate than ♂ and 2–3 mm. long; corolla as ♂ but lobes generally more thickened; anthers ± 0.5 mm. long, with little or no pollen; style caducous.
    Male
    Functionally ♂ flowers borne in 3–20-flowered, sessile or rarely pedunculate (up to 2 mm.) umbels; pedicels 3–7 mm. long, glabrous or sometimes pubescent; calyx-tube reduced; limb lobed almost to base; lobes somewhat unequal, linear-triangular, occasionally slightly spathulate, 0.75–2 mm. long, often with a few hairs at apex; corolla whitish or yellowish green, readily caducous; tube ± 1.5 mm. long, with a ring of deflexed hairs at throat inside; lobes triangular-ovate, 1.25–1.5 mm. long, 1–1.5 mm. wide at base, acute, often with a few hairs at apex outside, erect; anthers exserted but almost sessile, 0.75 mm. long; style shortly exceeding corolla-tube; stigmatic knob 0.25 mm. long and wide.
    Female
    Functionally ♀ flowers solitary; pedicels 0.4–1 cm. long; calyx-tube 1.75–2.5 mm. long; lobes more strongly spathulate than ♂ and 2–3 mm. long; corolla as ♂ but lobes generally more thickened; anthers ± 0.5 mm. long, with little or no pollen; style caducous.
    Fruits
    Fruit dark maroon, laterally compressed obovoid, 0.7–1.2 cm. long, 7–9 mm. wide, not indented at apex; calyx-lobes persistent; pedicel lengthening to 2 cm.; pyrene obovoid, flattened on ventral face, 7 mm. long, 3.5 mm. wide, crustaceous, surface ± smooth except for shallow crest around apex.
    Habitat
    Coastal bushland and wooded grassland to forest; 0–1200 m.
    Distribution
    K4 K7 P T3 T6 T8 Z
    [FZ]

    Rubiaceae, D. Bridson. Flora Zambesiaca 5:2. 1998

    Habit
    Compact, occasionally straggling, shrub or small tree 2–7 m tall (rarely a liana); bark pale grey, glabrous, or occasionally pubescent.
    Leaves
    Leaf blades pale greyish to bronze above when dry, 2.2–13 × 1–7.5 cm, broadly elliptic to round or broadly oblong-elliptic, obtuse to rounded at apex, obtuse to rounded or sometimes truncate at base, often unequal, glabrous or sometimes pubescent beneath; lateral nerves in 4 main pairs; tertiary nerves always obscure; domatia absent or inconspicuous; petiole 2–10 mm long; stipules light brown when dry, 4–16 × 2–10 mm, ovate or sometimes narrowly ovate, acuminate.
    Flowers
    Flowers 5–6-merous: functionally male flowers borne in 3–20-flowered, sessile or rarely shortly pedunculate (peduncle up to 2 mm long) umbels; functionally female flowers solitary; pedicels glabrous or sometimes pubescent, 3–7 mm long in functionally male flowers and 4–10 mm long in functionally female flowers.
    Calyx
    Calyx tube reduced in functionally male flowers, 1.75–2.5 mm long in functionally female flowers; calyx limb lobed almost to the base; lobes in functionally male flowers somewhat unequal, 0.75–2 mm long, linear-triangular, occasionally slightly spathulate; lobes in functionally female flowers usually broader, 2–3 mm long, more strongly spathulate and often with a few hairs at the apex.
    Corolla
    Corolla whitish or yellowish-green, readily caducous; tube c. 1.5 mm long with a ring of deflexed hairs at the throat inside; lobes 1.25–1.5 × 1–1.5 mm wide at base, triangular-ovate, acute, often with a few hairs at the apex outside, erect, apex generally more thickened in functionally female flowers than in functionally male flowers.
    Anthers
    Anthers present in both flower forms, exserted but almost sessile, c. 0.75 mm long in functionally male flowers, c. 0.5 mm long in functionally female flowers.
    Style
    Style shortly exceeding corolla tube.
    Pollen presenter
    Pollen presenter 0.25 mm long and wide, persistent in functionally male flowers and caducous in functionally female flowers.
    Fruits
    Fruit dark maroon when mature, 7–12 × 7–9 mm, flattened obovoid, not indented at the apex; calyx lobes persistent; pedicel lengthening to 20 mm.
    Pyrene
    Pyrene 7 × 3.5 mm, obovoid, with ventral face flattened, crustaceous, surface ± smooth except for a shallow crest around the apex.

    Images

    Distribution

    Native to:

    Kenya, Mozambique, Somalia, Tanzania

    Bullockia mombazensis (Baill.) Razafim., Lantz & B.Bremer appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Greenway, P.J. [2179], Tanzania K000412408 Unknown type material
    Greenway, P.J. [2179], Tanzania K000412409 Unknown type material
    Greenway, P.J. [2179], Tanzania K000412410
    Scott Elliot, G.F. [6380], Tanzania K000412411 Unknown type material
    Boivin, M. [s.n.], Tanzania K000172760 isotype

    First published in Ann. Missouri Bot. Gard. 96: 175 (2009)

    Accepted by

    • Govaerts, R.H.A. (2011). World checklist of selected plant families published update Facilitated by the Trustees of the Royal Botanic Gardens, Kew.

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0