1. Family: Arecaceae Bercht. & J.Presl
    1. Genus: Calamus L.
      1. Calamus hallierianus (Becc. ex K.Heyne) W.J.Baker

        This species is accepted, and its native range is W. Sumatera, Borneo.

    [PW]
    General Description
    Moderate-sized clustering rattan rarely exceeding 15 m in length. Stems closely clustering by short suckers, 6-11 mm diam. without sheaths, to 30 mm with sheaths; internodes 20-35 cm long, occasionally still longer in juvenile plants. Leaf sheath pinky-grey-green when fresh and newly emerged, dull grey-green when fresh and old, dark grey-brown when dry, densely covered with caducous scurfy grey indumentum (dark brown scales rather rare), and armed with groups and partial whorls of easily detached dark brown, spiculate spines to 9 cm long, borne on pale yellowish-green bulbous bases, these last remaining long after the spiculae have become detached; occa- sionally with broad, thin, almost papery, straw-coloured spines to 3.7 x 0.3 m long in groups among the spiculae, very rarely these spines abundant and spiculae almost absent; geniculus prominent, either armed with spiculae or, in forms with broad papery spines, almost inerm; ocrea to 1.5 cm, brown, tattering. Leaf 1.5-2.5 m long in adult; petiole 10-35 cm long, occasionally longer in juvenile leaves, to 10 mm wide and 7 mm thick, rounded abaxially, flattened or slightly grooved adaxially, armed with scattered spiculae below, and groups of 2-4 more robust, not easily-detached yellow-green spines to 1.1 cm long, especially along the margins, and scattered smaller spines on the adaxial face; minute papillae and caducous grey-white indumentum between the spines; petiole decreasing in width and armature above; rachis armed as the petiole but in upper part, spines tending to be confined to abaxial face and grouped in grapnels of 3.5; cirrus to 1.5 m long, with grapnel spine groups 5 cm distant below, decreasing above, and scattered grey-white indumentum and few chocolate-coloured scales. Leaflets 7-8 on each side of the rachis in mature leaf, the middle ones larger than the others, the largest broadly cuneate-rhomboid, to 35 x 16 cm at c. 20 cm from the insertion on the rachis; lowermost leaflets on either side c. 28 x 6 cm, upper- most to 32 x 10 cm; upper leaflet margins praemorse with rather rounded lobes, armed with small black bristles, the central lobe often prominent; whole leaflet markedly plicate with up to 15 plications, dull mid-green above, gleaming grey-white indumentose below, scattered dark-grey indumentum and brown pseudoparenchymatous scales along the main ribs on the upper surface and along margins exposed in bud; transverse veinlets rather obscure; emerging leaflets pink, turning dull brown then green. Staminate and pistillate inflorescences superficially similar, sessile, erect, very robust, the pistillate to 80 x 9 cm; base of inflorescence at point of insertion on leaf sheath c. 15 mm wide by 5 mm through. Prophyll pale pinky-green at first emergence from sheaths, then drying to dull brown, very stiff, almost woody in texture, with dense caducous chocolate-coloured indumentum on both surfaces but especially on the abaxial surface; beak to 8 cm long by 2 cm wide at base and 0-5 cm wide at tip. Axis branching to 2 orders with up to 8 first-order branches 8 cm long; first-order bracts tightly sheathing, to 8 mm long, with apiculus to 12 mm long. Staminate flowers solitary, borne on an alveolus c. 0.75 mm diam.; calyx with shallowly triangular lobes to 0.75 mm long; corolla c. 4.5 mm long split ? to the base into 3 lobes 2.25 mm wide near the base, tapering to bluntly triangular tips; stamens 6; filaments c. 1 mm long; anthers 1.75 x 0.2 mm; pollen yellow; pistillode trifid, minute. Pistillate flower borne with sterile staminate flower in a pair on an alveolus c. 1.25 mm diam.; calyx with 3 triangular lobes c. 1 mm high; corolla split to I into 3 broadly triangular lobes 4.5 x 3 mm; staminodes 6, with broad triangular bases and empty anthers c. 1.5 mm long; ovary c. 2.75 mm diam., rounded, tipped with 3 sinuous stigmas to 1-5 mm long. Sterile staminate flower to 6.5 mm long; calyx 1 mm long, shallowly lobed; corolla split for 4.5 mm; staminodes with filaments 1.75 mm long and anthers 1.5 x 0.2 mm. Mature fruit bursting the enclosing inflorescence bract, and tending to hang out, rounded, to 1.5 cm diam, tipped by stigma remains and covered with 12 vertical rows of pale orange-brown scales with darker brown margins; seed rounded c. 1.2 cm diam., covered in a thin sarcotesta; endosperm shallowly ruminate; embryo basal. Germination adjacent- ligular; eophyll not known.
    Distribution
    Widespread but rather local, possibly overlooked, in Sarawak, West and South Kalimantan, and Sabah, and one population in West Sumatra. Confined to lowland Mixed-Dipterocarp forest, sometimes on seasonally flooded alluvial flats.

    Distribution

    Native to:

    Borneo, Sumatera

    Calamus hallierianus (Becc. ex K.Heyne) W.J.Baker appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Apr 14, 2005 Watanabe, N.M. [NMW20], Indonesia Ceratolobus discolor K000209415
    Apr 14, 2005 Watanabe, N.M. [NMW38], Brunei Ceratolobus discolor K000209429
    Jul 21, 2000 Niga [BRUN15120], Brunei Ceratolobus discolor K000113142
    Jun 26, 1990 Wong, K.M. [489], Brunei Ceratolobus discolor K000525425
    Sep 16, 1977 Ramos [1490], Malaysia Ceratolobus discolor K000521016
    Sep 16, 1977 Ramos [1490], Malaysia Ceratolobus discolor K000521017
    Sep 16, 1977 Ramos [1490], Malaysia Ceratolobus discolor K000521018
    Sep 16, 1976 Dransfield, J. [JD3961], Indonesia Ceratolobus discolor K000521029
    Sep 16, 1976 Dransfield, J. [3986], Indonesia Ceratolobus discolor K000521028
    Sep 16, 1976 Dransfield, J. [JD3987], Indonesia Ceratolobus discolor K000525421
    Ashton, P.S. [12138], Malaysia Ceratolobus discolor K000521014
    Ashton, P.S. [12138], Malaysia Ceratolobus discolor K000521015
    Coode, M. [6830], Brunei Ceratolobus discolor K000525426
    Whitmore, T. [3130], Malaysia Ceratolobus discolor K000521024
    Mogea, J.P. [4421], Indonesia Ceratolobus discolor K000525422
    Mogea, J.P. [7398], Indonesia Ceratolobus discolor K000030612
    Dransfield, J. [4634], Malaysia Ceratolobus discolor K000521027
    Dransfield, J. [4663], Malaysia Ceratolobus discolor K000521023
    Dransfield, J. [4628], Malaysia Ceratolobus discolor K000521022
    Dransfield, J. [5972], Malaysia Ceratolobus discolor K000521021
    Dransfield, J. [6077], Malaysia Ceratolobus discolor K000521020
    Dransfield, J. [5275], Malaysia Ceratolobus discolor K000521019
    Dransfield, J. [4717], Malaysia Ceratolobus discolor K000525428
    Dransfield, J. [4644], Malaysia Ceratolobus discolor K000525427
    Christensen [1421], Malaysia Ceratolobus discolor K000521026
    Andersen, J. [13279], Malaysia Ceratolobus discolor K000521012
    Andersen, J. [13279], Malaysia Ceratolobus discolor K000521013
    Andersen, J. [13279], Malaysia Ceratolobus discolor K000521025
    Sambuling [92], Malaysia Ceratolobus discolor K000525423
    Tadong [722], Malaysia Ceratolobus discolor K000525424

    First published in Phytotaxa 204: 235 (2015)

    Sources

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Palmweb - Palms of the World Online
    Palmweb 2011. Palmweb: Palms of the World Online. Published on the internet http://www.palmweb.org. Accessed on 21/04/2013
    Content licensed under Creative Commons Attribution-NonCommercial-ShareAlike 3.0 Unported License http://creativecommons.org/licenses/by-nc-sa/3.0