1. Family: Rutaceae Juss.
    1. Genus: Zanthoxylum L.
      1. Zanthoxylum trijugum (Dunkley) P.G.Waterman

        This species is accepted, and its native range is Central Tanzania to Zambia.

    [FTEA]

    Rutaceae, J.O. Kokwaro (University of Nairobi). Flora of Tropical East Africa. 1982

    Habit
    Shrub or small tree usually 2–3 m. high, sometimes several-stemmed; bark grey, with scattered straight or slightly recurved spines and prickles 2–8 mm. long.
    Leaves
    Leaves 15–35 cm. long, rusty pubescent or greyish tomentose to glabrescent; rhachis sometimes sparsely armed; petiolules 0-1(–1.5) mm. long; leaflets 3–4 pairs, opposite or subopposite, ovate to elliptic, the first pair more ovate to deltoid, 4–12.2 cm. long, 2.5–6 cm. broad, acute to acuminate at the apex, broadly cuneate to shallowly cordate at the base, obscurely crenate-serrate; lateral nerves 6–10 pairs.
    Inflorescences
    Inflorescence a terminal pyramidal panicle 10–18 cm. long; peduncle pubescent; flowers clustered, on pedicels 0.5–1 mm. long.
    Calyx
    Sepals 4, slightly united at the base, ovate to subcircular, 0.6–1 mm. long, ciliolate.
    Corolla
    Petals 4, 1.5–3.5 mm. long, 1–1.3 mm. broad.
    Male
    Male flowers:stamens 4, 2–3 mm. long; gynophore pulviniform, glabrous; vestigial ovary obsoletely lobed.
    Flowers
    Female flowers not seen. Male flowers:stamens 4, 2–3 mm. long; gynophore pulviniform, glabrous; vestigial ovary obsoletely lobed.
    Female
    Female flowers not seen.
    Fruits
    Fruit globose, 4.5–6 mm. in diameter, ripening brown, reticulate and densely gland-dotted.
    Seeds
    Seed subglobose, 4.5–5.5 mm. in diameter, glossy black.
    Habitat
    Deciduous woodland, bushland and thicket, sometimes riverine; 1100–1450 m.
    Distribution
    T4 T5
    [FZ]

    Rutaceae, F. A. Mendoça. Flora Zambesiaca 2:1. 1963

    Habit
    Shrub or small tree; branches, leaves and inflorescences rusty-pubescent or greyish-tomentellous, glabrescent; stems and leaf-rhachis ± sparsely aculeate with straight or slightly recurved aculei 2–8 mm. long, rhachis of leaves sometimes unarmed.
    Leaves
    Leaves 15–35 cm. long; leaflets papyraceous, 3–4-jugate, opposite or subopposite, sessile or petiolulate with petiolules up to 1·5 mm. long, the terminal one with petiolule up to 30 mm. long, obtuse or rounded to shallowly cordate at the base; lamina 4–10 × 3–5·5 cm., ovate to elliptic, densely dotted with pellucid glands, acute at the apex, margin crenulate to minutely serrate, obtuse or rounded at the base; lateral nerves 6–10 pairs.
    Inflorescences
    Inflorescence a terminal panicle 12–18 cm. long.
    Flowers
    Flowers usually in cymose clusters with pedicels up to 1 mm. long; buds 3 mm. long, ellipsoid. Female flowers: not seen. Male flowers: stamens 4; filaments linear, bent inwards near the apex in bud; anthers dorsifixed, deeply 2-lobed at the base, laterally dehiscent; disk or gynophore pulviniform, glabrous; vestigial ovary obsoletely lobed.
    Calyx
    Sepals 4, 1 mm. long, slightly united at the base, ovate to subcircular, imbricate, persistent, margin ciliolate.
    Corolla
    Petals 4, 3·5 × 1·3 mm., imbricate.
    Male
    Male flowers: stamens 4; filaments linear, bent inwards near the apex in bud; anthers dorsifixed, deeply 2-lobed at the base, laterally dehiscent; disk or gynophore pulviniform, glabrous; vestigial ovary obsoletely lobed.
    Female
    Female flowers: not seen.
    Fruits
    Fruit 5 mm. in diam., globose, densely gland-dotted.
    Seeds
    Seed subglobose, black.

    Images

    Distribution

    Native to:

    Tanzania, Zambia, Zaïre

    Zanthoxylum trijugum (Dunkley) P.G.Waterman appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Martin, J.D. [137], Zambia K000199405 Unknown type material
    Martin, J.D. [493] K000199407 Unknown type material
    Martin, J.D. [571], Zambia K000199406 Unknown type material

    First published in Taxon 23: 365 (1975)

    Accepted by

    • Kokwaro, J.O. (1982). Flora of Tropical East Africa, Rutaceae: 1-52.

    Literature

    Kew Backbone Distributions
    • Exell, A.W., Fernandes, A. & Wild, H. (eds.) (1963). Flora Zambesiaca 2(1): 1-351. Royal Botanic Gardens, Kew.
    Flora of Tropical East Africa
    • Waterm. in Taxon 24: 365 (1975).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0