1. Family: Santalaceae R.Br.
    1. Genus: Thesium L.
      1. Thesium bangweolense R.E.Fr.

        This species is accepted, and its native range is N. Zambia to N. Malawi.

    [FZ]

    Flora Zambesiaca. Vol. 9, Part 3. Polygonaceae-Myriaceae. Pope GV, Polhill RM, Martins ES. 2006.

    Type
    Type:Zambia, Kapata–Bangweulu (Bangweolo) Peninsula, Kamindas [11°41'S 29°43'E], fl. & fr. 5.x.1911, R.E. Fries 890 (UPS, holotype; K).
    Habit
    Perennial herb drying yellowish-green, with short stout woody caudex c. 12 mm in diameter crowned with many very short closely scaly shoots eventually elongating into slender erect stems up to c. 350 mm tall, c. 1 mm diameter at base, simple to well branched near base, ± virgately branched above, branches sharply ascending, stems shallowly ribbed by decurrent scales, closely bracteate near base, more distantly so above Perennial herb drying yellowish-green, with short stout woody caudex c. 12 mm in diameter crowned with many very short closely scaly shoots eventually elongating into slender erect stems up to c. 350 mm tall, c. 1 mm diameter at base, simple to well branched near base, ± virgately branched above, branches sharply ascending, stems shallowly ribbed by decurrent scales, closely bracteate near base, more distantly so above.
    Style
    Style 1.2–2 mm long; stigma reaching apex of hood (well above anthers)
    Fruits
    Fruit 2.5 × 2 mm, pallid, strongly ribbed and reticulate. Fruit 2.5 × 2 mm, pallid, strongly ribbed and reticulate.
    Ecology
    Tall grass in wet meadows (Zambia), montane grassland (Malawi).
    Note
    The description of T. bangweolense was drawn up from the type specimen only. It is a distinctive plant differing from the other species with flowers subtended by an involucre of bracts by the combination of very slender stems ± virgately branched, and spiciform inflorescences with flowers solitary at the tips of very short scaly brachyblasts, the whole plant drying yellow-green. The specimen from Malawi, cited above, is in first flower, and shows spiciform inflorescences similar to those of T. bangweolense, but the plant is too young to judge what the final growth form will be. Thesium bangweolense remains a thoroughly ill-known species.
    Distribution
    Known with certainty only from Zambia; the Malawian specimen is cited with reservation. Malawi ZAM N, MAL N Zambia
    Leaves
    Leaves all reduced to sharply ascending scales, those near bases of stems crowded, c. 2.4 × 1.6 mm, distant upwards, c. 1.5–2 × 0.6–0.8 mm, lanceolate, cuspidate, thick-textured, margins membranous, minutely erose-ciliate. Leaves all reduced to sharply ascending scales, those near bases of stems crowded, c. 2.4 × 1.6 mm, distant upwards, c. 1.5–2 × 0.6–0.8 mm, lanceolate, cuspidate, thick-textured, margins membranous, minutely erose-ciliate
    Flowers
    Flowers solitary, sessile at tips of very short (c. 1–5(10) mm at extreme base), scaly brachyblasts arranged in very long slender lax spiciform inflorescences accounting for most of the stem's length. Flowers solitary, sessile at tips of very short (c. 1–5(10) mm at extreme base), scaly brachyblasts arranged in very long slender lax spiciform inflorescences accounting for most of the stem's length
    Bracts
    Bracts c. 6 forming an involucre below the flower, c. 1.5–2 × 0.8–1 mm, broadly lanceolate, cuspidate, thick-textured, margins membranous, minutely erose-ciliate, apices often rufous, whitish with age. Bracts c. 6 forming an involucre below the flower, c. 1.5–2 × 0.8–1 mm, broadly lanceolate, cuspidate, thick-textured, margins membranous, minutely erose-ciliate, apices often rufous, whitish with age
    Pedicel
    Pedicels wanting. Pedicels wanting
    Perianth
    Perianth white inside, yellowish-green outside; tube c. 0.5–0.8 mm long, no external glands visible; lobes c. 1.6–2 × 0.7 mm at base, oblong, obtuse, hood c. 0.3–0.6 mm long, margins inflexed, usually somewhat erose, sometimes with a few fimbrils on lower part. Perianth white inside, yellowish-green outside; tube c. 0.5–0.8 mm long, no external glands visible; lobes c. 1.6–2 × 0.7 mm at base, oblong, obtuse, hood c. 0.3–0.6 mm long, margins inflexed, usually somewhat erose, sometimes with a few fimbrils on lower part
    Stamens
    Stamens inserted at base of perianth lobes; filaments 0.2–0.5 mm long, almost hidden by anthers; anthers 0.4–0.8 mm long. Stamens inserted at base of perianth lobes; filaments 0.2–0.5 mm long, almost hidden by anthers; anthers 0.4–0.8 mm long
    Pistil
    Style 1.2–2 mm long; stigma reaching apex of hood (well above anthers).

    Images

    Distribution

    Native to:

    Malawi, Zambia

    Thesium bangweolense R.E.Fr. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Aug 23, 1949 Fries, R.E. [890], Zambia K000431508 Unknown type material

    First published in Wiss. Erg. Schwed. Rhodesia-Kongo-Exp. 1911-1912, 1: 21 (1914)

    Accepted by

    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • Wiss. Ergebn. Schwed. Rhod.-Kongo-Exped.: 21, fig. 2 (1914). TAB. 47, fig. C.
    Kew Backbone Distributions
    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0