1. Family: Santalaceae R.Br.
    1. Genus: Thesium L.
      1. Thesium fimbriatum A.W.Hill

        This species is accepted, and its native range is W. & S. Tanzania to S. Tropical Africa.

    [FZ]

    Flora Zambesiaca. Vol. 9, Part 3. Polygonaceae-Myriaceae. Pope GV, Polhill RM, Martins ES. 2006.

    Type
    Type:Malawi, between Khondowe and Karonga, Whyte 321 p.p. (K, holotype).
    Habit
    Possibly annual or perennial; stem 270–850 mm tall, (2.5)3–4 mm diameter at base, usually solitary, rarely 2–3 from woody crown, either simple or simple near base, soon virgately branched, the branches sharply ascending, strongly ribbed by raised vascular strands terminating in midribs of scale leaves and bracts, distantly leafy
    Leaves
    Leaves all reduced to scales, c. 0.5–2 × 0.4–1.6 mm, increasing in size upwards, distant below, becoming more crowded upwards and passing into floral bracts, lanceolate, acuminate, closely appressed, midrib strongly raised on dorsal surface
    Flowers
    Flowers sessile in axil of each bract, up to c. 10 crowded on a very short brachyblast, these either simple or branched, or sometimes in long many-flowered spikes, the flowers ± crowded below, distant above, both brachyblasts and long spikes sometimes present on one plant, the whole forming a panicle
    Bracts
    Bracts (lowermost) 2.4–3.5 × 1–2 mm, ovate-acuminate, margins membranous, minutely erose-ciliate, keeled by strongly raised midrib; bracteoles 2, similar to bracts
    Pedicel
    Pedicels wanting
    Perianth
    Perianth possibly creamy-yellow inside; tube 0.5–1 mm long, 5 tiny (c. 0.1 mm) tooth-like glands alternating with the lobes; lobes usually 5, sometimes 4, c. 1.5–2.7 × 0.4–0.6 mm at base, subspathulate, apex subobtuse to acute, hood c. 0.25(0.4) mm long, margins inflexed, fringed with long white hairs
    Stamens
    Stamens inserted near base of perianth lobes; filaments conspicuous, c. 0.9–1.5 mm long, lower part adnate to perianth lobe; anthers 0.3–0.5 mm long
    Style
    Style c. 1.2–2.6 mm long; stigma reaching ± to middle of anthers
    Fruits
    Fruit 1.6–2 × 1.4–2 mm, buff-coloured to light reddish-brown, strongly ribbed, the 5 intermediate ribs often forking once near base, reticulations few and often obscure.
    Ecology
    Moist dambos or drier areas in grassland; c. 1370–1800 m.
    Note
    There are 4 specimens on the type sheet of T. fimbriatum. Hill segregated one of them as the type of the name and assigned the rest to T. stuhlmannii Engl. (type from Itiolo [misprint for Itolio in Bukoba Distr.] in NW Tanzania, west of Victoria Nyanza). The characters given to distinguish T. fimbriatum from T. stuhlmannii were its more bushy habit, more delicate branches, ovate acuminate ciliate bracts and linear elliptic perianth lobes, but these characters, apart from the delicate branches, apply equally well to the type specimen of T. stuhlmannii, which does, however, differ in its glabrous perianth lobes and tiny fruits (c. 1.2 × 1.7 mm). Whyte's three specimens assigned by Hill to T. stuhlmannii have the fimbriate lobes and slightly larger fruits of T. fimbriatum. Apparent differences in branching can be attributed to degree of development of brachyblasts, which may depend on the age of the plant: this needs field observation. Indeed, this whole difficult group (that is, those species with ovate acuminate bracts, clusters of sessile flowers, and tooth-like glands alternating with the perianth lobes, 5 species in the Flora Zambesiaca area, but many more in Angola, Dem. Rep. Congo and East Africa) needs study over a wide range of material. The type of T. shabense precisely matches that of T. fimbriatum. Both specimens often have the perianth lobes slightly longer than the other material included by me under T. fimbriatum (lobes up to c. 1.8–3 mm versus 1.4–2.2 mm) but all the material agrees in its relatively stout stems, often solitary, always simple below, branching above, fimbriate perianth lobes with only a small hood, and tiny (c. 0.1 mm) external glands. T. fimbriatum A.W. Hill in F.C. 5, 2: 191 (1915) is a later homonym, renamed T. brachystylum A.W. Hill in 1920.
    Distribution
    Mozambique Malawi ZAM N, ZAM W, ZAM C, MAL N, MOZ N Also in Katanga (Shaba) Province of Dem. Rep. Congo and Tanzania. Zambia

    Images

    Distribution

    Native to:

    Malawi, Mozambique, Tanzania, Zambia, Zaïre

    Thesium fimbriatum A.W.Hill appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    K000431484
    Whyte, A. [321], Malawi K000431483 Unknown type material

    First published in Bull. Misc. Inform. Kew 1910: 184 (1910)

    Accepted by

    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.
    • Polhill, R.M. (2005). Flora of Tropical East Africa, Santalaceae: 1-27.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.

    Literature

    Flora Zambesiaca
    • Bull. Misc. Inform., Kew 1915: 27 (1915).
    • F.T.A. 6, 1: 422 (1911). TAB. 46, fig. D.
    Kew Backbone Distributions
    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0