1. Family: Santalaceae R.Br.
    1. Genus: Thesium L.
      1. Thesium schliebenii Pilg.

        This species is accepted, and its native range is SW. Tanzania to N. Zambia.

    [FTEA]

    Santalaceae, P.M. Polhill, B.A., Ph.D., F.L.S. Flora of Tropical East Africa. 2005

    Habit
    Perennial herb, with narrow woody rhizomes up to 5 cm long; stems several from each rhizome, the whole forming a many-stemmed tuft; stems erect, 25–60 cm tall, simple to virgately branched, flowering branches sharply ascending, later extending into sterile scaly tips; ribs strongly raised and extending into midribs of scale leaves and bracts.
    Leaves
    Leaves all reduced to scales, closely appressed, lanceolate, 1–2.5x0.4–1.2 mm, increasing in size upwards and passing into floral bracts, acuminate, midrib strongly raised on outer surface.
    Flowers
    Flowers sessile, with a bract and 2 similar bracteoles or an involucre with 3–5 bracts, usually up to 6 flowers crowded on very condensed short shoots, but sometimes also spaced on elongate shoots, both types occurring on the same plant; bracts and bracteoles ovate-acuminate, 2–3x1–1.4 mm, margins membranous, minutely erose-ciliate, keeled by strongly raised midrib.
    Perianth
    Perianth greenish white or yellow; tube 1 mm long, 5 tiny (up to 0.1 mm) tooth-like glands alternating with the lobes; lobes linear, 3–3.6x0.4–0.5 mm, tapered to a long hood, margins inflexed, erose in upper part, densely white-bearded in lower part; hood 0.6–0.8 mm long.
    Stamens
    Stamens inserted near base of perianth lobes; filaments conspicuous, 1.5–2.2 mm long, shortly adnate to lobe; anthers 0.4–0.7 mm long.
    Pistil
    Style 2.8–3.5 mm long; stigma reaching base of hood.
    Fruits
    Fruit buff-coloured to pale brown, globose, 2 mm in diameter, very strongly ribbed, very weakly reticulate.
    Habitat
    Wet grassland or swamps; 1600–2100 m
    Distribution
    T7
    [FZ]

    Flora Zambesiaca. Vol. 9, Part 3. Polygonaceae-Myriaceae. Pope GV, Polhill RM, Martins ES. 2006.

    Type
    Type from Tanzania (Lupembe area, Njombe Distr.).
    Stamens
    Stamens inserted at base of perianth lobes; filaments conspicuous, 1.5–2.2 mm long, briefly adnate to lobe; anthers 0.4–0.7 mm long Stamens inserted at base of perianth lobes; filaments conspicuous, 1.5–2.2 mm long, briefly adnate to lobe; anthers 0.4–0.7 mm long.
    Style
    Style 2.8–3.5 mm long; stigma reaching base of hood
    Fruits
    Fruit 2 × 2 mm, probably buff-coloured when fully ripe, very strongly ribbed, very weakly reticulate. Fruit 2 × 2 mm, probably buff-coloured when fully ripe, very strongly ribbed, very weakly reticulate.
    Ecology
    Dambo, swampy ground in long grass; 1525 m.
    Note
    Known to me only from the types of T. schliebenii and T. schaijesii and Mrs Richards' collection, cited above, which was found roughly midway between the two type localities. Mrs. Richards' specimens carry masses of old crowded brachyblasts from which the fruits have fallen, as well as younger shoots with lax spikes of flowers; both type specimens comprise younger stems without crowded brachyblasts, but the exceptionally long-hooded and very pointed perianth lobes are distinctive among species recorded in the region.
    Distribution
    Also in Tanzania and Dem. Rep. Congo. Zambia ZAM N
    Habit
    Perennial herb with woody stolons up to c. 50 mm long, stems several from each stolon, the whole forming a many-stemmed tuft; stems 230–550 mm long, 1–2.5 mm diameter at base, simple to laxly branched, branches sharply ascending, strongly ribbed by raised vascular strands terminating in midribs of scale leaves and bracts, distantly leafy. Perennial herb with woody stolons up to c. 50 mm long, stems several from each stolon, the whole forming a many-stemmed tuft; stems 230–550 mm long, 1–2.5 mm diameter at base, simple to laxly branched, branches sharply ascending, strongly ribbed by raised vascular strands terminating in midribs of scale leaves and bracts, distantly leafy
    Leaves
    Leaves all reduced to scales, c. 1–2.5 × 0.4–1.2 mm, increasing in size upwards and passing into floral bracts, lanceolate, acuminate, closely appressed, midrib strongly raised on dorsal surface. Leaves all reduced to scales, c. 1–2.5 × 0.4–1.2 mm, increasing in size upwards and passing into floral bracts, lanceolate, acuminate, closely appressed, midrib strongly raised on dorsal surface
    Flowers
    Flowers often in long lax spikes, sometimes solitary, usually up to 6 on very short brachyblasts, on older stems brachyblasts crowded on very short simple or branched side shoots, sometimes with long new shoots developing at their apices. Flowers often in long lax spikes, sometimes solitary, usually up to 6 on very short brachyblasts, on older stems brachyblasts crowded on very short simple or branched side shoots, sometimes with long new shoots developing at their apices
    Bracts
    Bracts and bracteoles 2–3 × 1–1.4 mm, ovate-acuminate, margins membranous, minutely erose-ciliate, keeled by strongly raised midrib, usually 3 only, rarely 4. Bracts and bracteoles 2–3 × 1–1.4 mm, ovate-acuminate, margins membranous, minutely erose-ciliate, keeled by strongly raised midrib, usually 3 only, rarely 4
    Pedicel
    Pedicels wanting. Pedicels wanting
    Perianth
    Perianth “weiss, aussen gelb” or “greeny white”; tube 1 mm long, 5 tiny (up to 0.1 mm) tooth-like glands alternating with the lobes; lobes 3–3.6 × 0.4–0.5 mm, linear, very acute, hood 0.6–0.8 mm long, margins inflexed, erose in upper part, densely white-bearded in lower. Perianth “weiss, aussen gelb” or “greeny white”; tube 1 mm long, 5 tiny (up to 0.1 mm) tooth-like glands alternating with the lobes; lobes 3–3.6 × 0.4–0.5 mm, linear, very acute, hood 0.6–0.8 mm long, margins inflexed, erose in upper part, densely white-bearded in lower
    Pistil
    Style 2.8–3.5 mm long; stigma reaching base of hood.

    Distribution

    Native to:

    Tanzania, Zambia, Zaïre

    Thesium schliebenii Pilg. appears in other Kew resources:

    First published in Notizbl. Bot. Gart. Berlin-Dahlem 11: 397 (1932)

    Accepted by

    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.
    • Polhill, R.M. (2005). Flora of Tropical East Africa, Santalaceae: 1-27.

    Literature

    Flora Zambesiaca
    • Notizbl. Bot. Gart. Berlin-Dahlem 11: 397 (1932).
    Kew Backbone Distributions
    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.
    Flora of Tropical East Africa
    • Pilg. in Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahelm 11: 397 (1932).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0