1. Family: Malvaceae Juss.
    1. Genus: Grewia L.
      1. Grewia flavescens Juss.

        This species is accepted, and its native range is Tropical & S. Africa to India.

    [FZ]

    Tiliaceae, H. Wild. Flora Zambesiaca 2:1. 1963

    Habit
    Shrub 2–5 m. tall; young branches stellate-pubescent, becoming glabrous; older stems 4-angled and somewhat sulcate, sometimes scandent.
    Leaves
    Leaf-lamina 4–12 × 2–8·5 cm., oblanceolate, obovate or oblong-lanceolate, occasionally almost circular, acute or acuminate at the apex, margins irregularly serrate, rounded or subcordate at the base, harshly pubescent, particularly below; petiole up to 7 mm. long but usually shorter, pilose; stipules up to 1 cm. long, subulate to oblong-lanceolate, somewhat keeled, pubescent on both sides.
    Inflorescences
    Inflorescences all axillary on pubescent peduncles up to 1·5 cm. long; pedicels up to 2 cm. long, 2–3 per peduncle, pubescent; bracts entire, c. 3 mm. long, lanceolate to ovate-lanceolate, pubescent, especially on the back.
    Buds
    Flower-buds oblong with a slight swelling and then a constriction just above the base.
    Calyx
    Sepals yellow inside, yellowish-tomentose outside, 12–20 mm. long, lorate.
    Corolla
    Petals yellow, 6–8 × 1·5 mm., narrowly oblong, with a nectariferous claw villous on the margins within and at the base outside.
    Androgynophore
    Androgynophore 1·5–2 mm. long, glabrous, not elongated above the node.
    Pistil
    Ovary closely setose-pubescent, entire or slightly 2-lobed; style c. 1 cm. long, glabrous; stigma-lobes usually 4, subulate.
    Fruits
    Fruit 8–14 × 13–15 mm., depressed-globose, sometimes entire but usually shallowly 2- or occasionally 4-lobed, yellowish-brown with short, appressed-stellate hairs often mixed with longer stellate hairs, somewhat shining when ripe.
    [FTEA]

    Tiliaceae, C. Whitehouse, M. Cheek, S. Andrews & B. Verdcourt. Flora of Tropical East Africa. 2001

    Habit
    Shrub or scandent tree to 7.5 m tall; young stems densely roughly to softly pubescent, often flattened, older branches 3–4-angled, grey-brown.
    Leaves
    Leaves broadlyelliptic, ovate, oblong, obovate or almost circular, 4.5–14.5 cm long, 2–11 cm wide, obtuse, acute or acuminate at the apex, rounded, truncate or slightly cordate at the base, usually scabrid above, roughly to softly densely pubescent beneath, sometimes ± tomentose; petiole 3–12 mm long; stipules 9–10 mm long.
    Inflorescences
    Inflorescence a 1–3-flowered cyme, 1–4 congested in a leaf-axil; peduncle 3–9 mm long; pedicels 5–14 mm long.
    Flowers
    Flowers yellow; sepals 7–22 mm long, green to reddish brown on the outside, yellow to orange inside; petals 8–15 mm long, yellow.
    Androgynophore
    Androgynophore 1–2 mm long, glabrous.
    Stamens
    Stamens 8–18 mm long, yellow.
    Pistil
    Ovary ± 3 mm long; style 4–17 mm long, yellow, hairy.
    Fruits
    Fruit shallowly or obscurely 1–3(–4)-lobed, 8–10 mm long, 7–14 mm wide, sparsely stellate-pubescent to glabrescent and shiny, green becoming orange to reddish brown, later brownish black.
    Figures
    Fig. 3/24 (leaf, p. 9).
    Habitat
    Commiphora woodland, thicket and bushland, often riverine or on rocky outcrops and termite mounds, sometimes scandent on the edge of forests; 600–1450 m
    Distribution
    K1 K2 T1 T2 T4 T5 T7 U1 tropical Africa south to Botswana, Namibia and NE South Africa
    [FWTA]

    Tiliaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:2. 1958

    Habit
    A savannah shrub
    Flowers
    Flowers yellow
    Fruits
    Ripe fruits brown.

    Images

    Distribution

    Native to:

    Angola, Botswana, Burkina, Burundi, Cameroon, Caprivi Strip, Central African Repu, Chad, Eritrea, Ethiopia, Gabon, Gambia, Guinea, India, Ivory Coast, Kenya, KwaZulu-Natal, Malawi, Mali, Mauritania, Mozambique, Namibia, Niger, Northern Provinces, Rwanda, Saudi Arabia, Senegal, Sudan, Swaziland, Tanzania, Togo, Uganda, Yemen, Zambia, Zaïre, Zimbabwe

    Common Names

    English
    Donkey berry, Rough-leaved raisin, Sandpaper raisin

    Grewia flavescens Juss. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Sep 16, 2008 Crawford, F. [64], Namibia K000450344
    Jan 1, 1958 Dinter [68], South Africa K000241916 syntype
    Jan 1, 1956 Kirk, J. [349], Mozambique K000975784
    Jan 1, 1956 Rautanen, M. [614] K000241929
    K000241712
    Heudelot [283] K000241713
    Timberlake, J. [s.n.], Malawi K000614741

    First published in Ann. Mus. Natl. Hist. Nat. 4: 91 (1804)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Meena, S.L. (2012). A checklist of the vascular plants of Banaskantha district, Gujarat, India Nelumbo 54: 39-91.
    • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Mannheimer, C.A. & Curtis, B.A. (eds.) (2009). Le Roux and Müller's field guide to the trees and shrubs of Namibia, rev. ed.: 1-525. Macmillan Education Namibia, Windhoek.
    • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Govaerts, R. (2003). World Checklist of Seed Plants Database in ACCESS G: 1-40325.
    • Whitehouse, C., Cheek, M., Andrews, S. & Verdcourt, B. (2001). Flora of Tropical East Africa, Tiliaceae & Muntingiaceae: 1-120.
    • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
    • Edwards, S., Tadesse, M. & Hedberg, I. (eds.) (1995). Flora of Ethiopia and Eritrea 2(2): 1-456. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
    • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
    • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 3: 1-89. ORSTROM, Bangui.
    • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Exell, A.W., Fernandes, A. & Wild, H. (eds.) (1963). Flora Zambesiaca 2(1): 1-351. Royal Botanic Gardens, Kew.

    Literature

    Flora of West Tropical Africa
    • Aubrév. Fl. For. Soud.-Guin. 154, t. 26, 6–7.
    • Burret in Engl. Bot. Jahrb. 45: 168
    • in Ann. Mus. Paris 4: 91 (1804)
    Kew Backbone Distributions
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Meena, S.L. (2012). A checklist of the vascular plants of Banaskantha district, Gujarat, India Nelumbo 54: 39-91.
    • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
    • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
    • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
    Flora of Tropical East Africa
    • Sebsebe in Fl. Eth. 2(2): 149, fig. 79.1/6–8 (1995).
    • Beentje, Kenya Trees, Shrubs and Lianas p. 154, fig. f4 on 152, map (1994).
    • Wild in Flora of Southern Africa 21: 4 (1984).
    • Wild & Gonç. in Fl. Moçamb. 28: 15 (1969).
    • R. Wilczek in Flore du Congo Belge et du Ruanda-Urundi, 10: 25 (1963).
    • Wild in Flora Zambesiaca 2: 46 (1963).
    • F. White, Forest Flora of Northern Rhodesia p. 239, fig. 42f (1962).
    • Dale & Greenway, Kenya Trees and Shrubs p. 565, fig. 103/f4 (1961).
    • Keay in Flora of West Tropical Africa, ed. 2, 1: 305 (1958).
    • Excell & Mendonça in Consp. Fl. Angol. 1: 221 (1951).
    • F. W. Andr., The Flowering Plants of the Anglo-Egyptian Sudan 1: 223 (1950).
    • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 618 (1949).
    • Burret in A. Engler, Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 45: 168 (1910).
    • Juss. in Ann. Mus. Hist. Nat. Paris 4: 91 (1804).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Plants and People Africa
    Common Names from Plants and People Africa http://www.plantsandpeopleafrica.com/
    © Plants and People Africa http://www.plantsandpeopleafrica.com http://creativecommons.org/licenses/by/3.0/