1. Family: Malvaceae Juss.
    1. Genus: Grewia L.
      1. Grewia villosa Willd.

        This species is accepted, and its native range is Africa to Indian Subcontinent.

    [FWTA]

    Tiliaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:2. 1958

    Habit
    A coarse-leaved shrub
    Ecology
    Of the drier savannah regions, often on rocky hills
    Height
    10–15 ft. high
    Flowers
    Flowers reddish-brown.
    [FTEA]

    Tiliaceae, C. Whitehouse, M. Cheek, S. Andrews & B. Verdcourt. Flora of Tropical East Africa. 2001

    Habit
    Shrub 1.2–4.5 m high, plants apparently either bisexual or unisexual; stems flattened; young shoots densely covered with villous hairs.
    Leaves
    Leaves broadly ovate to orbicular, 2–15 cm long, 1.6–18.5 cm wide, rounded to obtuse or slightly apiculate at the apex, rounded to cordate at the base, margin serrate, pubescent above, more so on the veins, densely stellate-pubescent beneath with long simple hairs on the veins; veins prominent and finely reticulate, leaves appearing bullate when young; petiole 0.6–5.5 cm long, densely villous; stipules broadly ovate to ovate-lanceolate, 5–10 mm long, caducous.
    Inflorescences
    Inflorescence a 2–10-flowered cyme.
    Flowers
    Flowers dull red, yellow or brownish; sepals 5–10 mm long, densely hairy on the outside, usually reddish on the inside; petals much smaller than sepals, 1–3 mm long, usually pinkish.
    Androgynophore
    Androgynophore glabrous.
    Stamens
    Stamens up to 3 mm long; filaments whitish; anthers yellow.
    Pistil
    Ovary densely villous; style 1.5–3 mm long; stigma much divided.
    Fruits
    Fruit unlobed, subglobose, flattened vertically, 1–1.6 cm in diameter, orange to rusty-brown, covered with small warts tipped by long hairs, outer wall crustaceous and well-separated from the pulp.
    Figures
    Fig. 6.
    Habitat
    Dry Acacia-Commiphora bushland, rocky hillsides and beside dry watercourses; 0–1550 m
    Distribution
    widespread throughout the drier parts of Africa from Senegal to Sudan and Ethiopia and south to South Africa K1 K2 K3 K4 K5 K6 K7 T2 T3 T5 T7 U1 U3
    [FZ]

    Tiliaceae, H. Wild. Flora Zambesiaca 2:1. 1963

    Androgynophore
    Androgynophore up to 0·75 mm. long, glabrous, with a more or less membranous undulate rim.
    Pistil
    Ovary not lobed, densely villous; style c. 3 mm. long, pubescent, especially towards the base; stigma divided into many laciniate segments.
    Fruits
    Fruit reddish, c. 15 mm. in diam., globose, shallowly 4-lobed, with small scattered tubercles bearing rather long caducous setulose hairs.
    Habit
    Much-branched shrub 1–3 m. tall; young branchlets with yellowish silky hairs.
    Leaves
    Leaf-lamina up to 12 cm. in diam., subcircular to broadly elliptic, apex rounded, margin serrate, slightly cordate at the base and sometimes asymmetric, sparsely pubescent, green and finely reticulate above, grey-pubescent or villous below; veins prominent and reticulate; petiole up to 4 cm. long, setulose-pubescent; stipules 5–12 mm. long, ovate to broadly oblanceolate, pubescent, submembranous.
    Inflorescences
    Inflorescences leaf-opposed or frequently arising some distance from the node; peduncle up to 5 mm. long, pubescent; pedicels similar, up to 3 mm. long; bracts c. 7 mm. long, ovate, similar to the stipules.
    Buds
    Flower-buds globose to cylindric-globose.
    Calyx
    Sepals 5–9 mm. long, linear-lanceolate, silky-pubescent outside, slightly pubescent and yellow inside.
    Corolla
    Petals yellow, about half the length of the sepals, oblong or obovate-oblong, with a basal nectariferous claw circumvillous within but not extended above into a narrow ledge, nectary and lamina of about equal length, pubescent behind the nectary and along the mid-line of the lamina.

    Images

    Distribution

    Native to:

    Angola, Bangladesh, Benin, Botswana, Burkina, Cameroon, Cape Verde, Caprivi Strip, Central African Repu, Chad, Djibouti, Eritrea, Ethiopia, Ghana, Gulf States, India, Iran, Ivory Coast, Kenya, KwaZulu-Natal, Mali, Mauritania, Mozambique, Namibia, Niger, Nigeria, Northern Provinces, Oman, Pakistan, Palestine, Saudi Arabia, Senegal, Somalia, Sudan, Swaziland, Tanzania, Togo, Uganda, Yemen, Zaïre, Zimbabwe

    Grewia villosa Willd. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Sep 16, 2008 Crawford, F. [28], Namibia K000450330
    Sep 16, 2008 Crawford, F. [04], Namibia K000450346
    Sep 16, 2008 Crawford, F. [11], Namibia K000450347
    Feb 23, 1909 Hooker, J.D. [173], Cape Verde K001134258
    Feb 23, 1909 Hooker, J.D. [173], Cape Verde K001134259
    Feb 23, 1909 Kotschy [386], Ethiopia K000241764
    Schimper [1278], Ethiopia K000241765
    Schimper [878], Ethiopia K000241766
    Kotschy [386], Ethiopia K000241767
    Kotschy [381], Sudan K000241768
    [Periottel], D. [s.n.], Senegal K000241811
    s.coll. [Cat. no. 6306] K001123543
    s.coll. [Cat. no. 6306], India K001123544

    First published in Neue Schriften Ges. Naturf. Freunde Berlin 4: 205 (1803)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Dobignard, A. & Chatelain, C. (2012). Index synonymique de la flore d'Afrique du nord 4: 1-431. Éditions des conservatoire et jardin botaniques, Genève.
    • Onana, J.M. (2011). The vascular plants of Cameroon a taxonomic checklist with IUCN assessments: 1-195. National herbarium of Cameroon, Yaoundé.
    • Mannheimer, C.A. & Curtis, B.A. (eds.) (2009). Le Roux and Müller's field guide to the trees and shrubs of Namibia, rev. ed.: 1-525. Macmillan Education Namibia, Windhoek.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Govaerts, R. (2003). World Checklist of Seed Plants Database in ACCESS G: 1-40325.
    • Whitehouse, C., Cheek, M., Andrews, S. & Verdcourt, B. (2001). Flora of Tropical East Africa, Tiliaceae & Muntingiaceae: 1-120.
    • Boulos, L. (2000). Flora of Egypt 2: 1-352. Al Hadara Publishing, Cairo.
    • Jongbloed, M., Western, R.A. & Boer, B. (2000). Annotated Check-list for plants in the U.A.E.: 1-90. Zodiac Publishing, Dubai.
    • Collenette, S. (1999). Wildflowers of Saudi Arabia: 1-799. National commission for wildlife conservation and development (NCWCD), Kingdom of Saudi Arabia.
    • Thulin, M. (ed.) in Thulin, M. (ed.) (1999). Flora of Somalia 2: 1-303. The Royal Botanic Gardens, Kew.
    • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
    • Edwards, S., Tadesse, M. & Hedberg, I. (eds.) (1995). Flora of Ethiopia and Eritrea 2(2): 1-456. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Ghazanfar, S.A. (1992). An Annotated Catalogue of the Vascular Plants of Oman and their Vernacular names Scripta Botanica Belgica 2: 1-153.
    • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
    • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Browicz, K. (1981). Flora Iranica 148: 1-15. Naturhistorisches Museums Wien.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 3: 1-89. ORSTROM, Bangui.
    • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Exell, A.W., Fernandes, A. & Wild, H. (eds.) (1963). Flora Zambesiaca 2(1): 1-351. Royal Botanic Gardens, Kew.
    • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.

    Literature

    Flora of West Tropical Africa
    • Ic. Pl. Afr. t. 5.
    • Aubrév. Fl. For. Soud.-Guin. 154, t. 25, 5
    • Burret in Notizbl. Bot. Gart. Berl. 9: 721
    • Burret in Engl. Bot. Jahrb. 45: 198
    • —F.T.A. 1: 249
    Kew Backbone Distributions
    • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
    • Onana, J.M. (2011). The vascular plants of Cameroon a taxonomic checklist with IUCN assessments: 1-195. National herbarium of Cameroon, Yaoundé.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Jongbloed, M., Western, R.A. & Boer, B. (2000). Annotated Check-list for plants in the U.A.E.: 1-90. Zodiac Publishing, Dubai.
    • Collenette, S. (1999). Wildflowers of Saudi Arabia: 1-799. National commission for wildlife conservation and development (NCWCD), Kingdom of Saudi Arabia.
    • Edwards, S., Tadesse, M. & Hedberg, I. (eds.) (1995). Flora of Ethiopia and Eritrea 2(2): 1-456. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
    • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 3: 1-89. ORSTROM, Bangui.
    • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.
    Flora of Tropical East Africa
    • Sebsebe in Fl. Eth. 2(2): 152, fig. 79.3 (1995).
    • Audru et al., Pl. Vasc. Rép. Djibouti 2: 223, t. on p. 224, fig. 82 (1994).
    • Beentje, Kenya Trees, Shrubs and Lianas p. 159, fig. b, b1 on 152, map (1994).
    • K.M. Matthew in Journal of the Linnean Society of London, Botany 113: 364, 376 (1993).
    • Blundell, Wild Flow. E. Afr.: 69, fig. 418 (1987).
    • Wild in Flora of Southern Africa 21: 4 (1984).
    • Kokwaro, Medic. Pl. E. Afr.: 214 (1976).
    • Wild & Gonç. in Fl. Moçamb. 28: 35 (1969).
    • R. Wilczek in Flore du Congo Belge et du Ruanda-Urundi, 10: 30 (1963).
    • Wild in Flora Zambesiaca 2: 62 (1963).
    • Dale & Greenway, Kenya Trees and Shrubs p. 571, fig. 103/b & b1 (1961).
    • Keay in Flora of West Tropical Africa, ed. 2, 1: 304, fig. 114 (1958).
    • Excell & Mendonça in Consp. Fl. Angol. 1: 223 (1951).
    • F. W. Andr., The Flowering Plants of the Anglo-Egyptian Sudan 1: 220, fig. 119 (1950).
    • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 614 (1949).
    • Battiscombe & Dale, Trees and Shrubs of Kenya Colony p. 40 (1936).
    • Burtt-Davy, Check-lists For. Trees & Shrubs Brit. Emp. 1, Uganda: 117 (1935).
    • Burret in A. Engler, Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 45: 198 (1910).
    • K. Schum. in Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete, Theile C: 264 (1895).
    • Mast. in Flora of Tropical Africa 1: 249 (1868).
    • Willd. in Ges. Nat. Fr. Berlin, Neue Schr. 4: 205 (1803).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
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    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

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