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This species is accepted, and its native range is Old World.

[FWTA]

Zygophyllaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:2. 1958

Morphology General Habit
A trailing herb, woody below, from pubescent to tomentose
Ecology
A pest of footpaths and fields.

[FZ]

Zygophyllaceae, E. Launert. Flora Zambesiaca 2:1. 1963

Morphology General Habit
Annual herb, extremely variable in habit, with spreading prostrate and usually decumbent branches, ± pubescent, villous or hispid, sometimes glabrescent in all vegetative parts; branches up to 2 m. long, sometimes woody at the base.
Morphology Leaves
Leaves unequal, the larger up to 6 cm. long (usually somewhat smaller) with up to 8 pairs of leaflets; the smaller up to 3·5 cm. long (usually much smaller) with up to 6 pairs of leaflets; leaflets 4–15 × 1·5–7 mm., rather oblique, oblong, oblong-lanceolate or lanceolate-ovate, acute or subobtuse, villous on both surfaces or nearly glabrous.
Morphology Leaves Stipules
Stipules up to 10 mm. long, linear or linear-lanceolate, acute.
Morphology Reproductive morphology Inflorescences Peduncles
Peduncle usually shorter than, or as long as the subtending leaf.
Morphology Reproductive morphology Flowers Calyx
Sepals 3–6 mm. long, linear-lanceolate, usually villous outside.
Morphology Reproductive morphology Flowers Corolla
Petals light yellow, 3–8 (12) mm. long, broadly cuneate, usually shorter than or as long as or sometimes up to 2·3 times the length of the sepals.
Morphology Reproductive morphology Flowers Androecium Stamens
Filaments about 3 mm. long; anthers 0·5–1 (1·2) mm. long.
Morphology Reproductive morphology Flowers Gynoecium Pistil
Style very short; stigma 0·8–1·2 mm. long, hemispheric, mostly asymmetric and nearly sessile on the ovary.
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary with stiff bulbous-based hairs.
Morphology General Glands
Intrastaminal glands not connate.
Morphology Reproductive morphology Fruits
Fruit breaking up into 5 cocci, each with 2 ± strong divergent spines in the upper and 2 smaller ones in the lower part, usually crested on the back, tuberculate and strigose.

[FSOM]

M. Thulin. Flora of Somalia, Vol. 1–4 [updated 2008] https://plants.jstor.org/collection/FLOS

Distribution
Tropical and subtropical regions throughout the world.
Morphology General Habit
Pubescent to villous annual-Leaves 2–6 cm long; leaflets 5–8 pairs, oblong to lanceolate, usually 4–18 x 1.5–5 mm; stipules linear to lanceolate, up to 8 mm long
Morphology Reproductive morphology Flowers Pedicel
Pedicel 5–10 mm long
Morphology Reproductive morphology Flowers Calyx
Sepals linear-lanceolate, 3–6 mm long
Morphology Reproductive morphology Flowers Corolla
Petals yellow, 4–8 mm long
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 8–10; filaments up to 3 mm long
Morphology Reproductive morphology Flowers Gynoecium Style
Style very short or absent; stigma ± pyramidal, c. 1 mm long
Morphology Reproductive morphology Fruits
Fruit disc-shaped; pedicel c. 10 mm long; mericarps 4–5 x 2–3 mm, with 2 pairs of spines.

[FTEA]

Zygophyllaceae, M. Nabil El Hadidi (Faculty of Science, Cairo University). Flora of Tropical East Africa. 1985

Morphology General Habit
Annual ± pubescent, villous or hispid herb; branches prostrate, up to 90 cm. long; internodes up to 6 cm. long, 2-3 mm. in diameter.
Morphology Leaves
Leaves 3-6 cm. long; leaflets 5-8 pairs, oblong to lanceolate, 4-8 × 1.5-5 mm.; stipules linear to lanceolate, up to 8 mm. long.
Morphology Reproductive morphology Flowers
Flower medium-sized, up to 15 mm. across; pedicel 5-10 mm. long, shorter than the subtending leaf.
Morphology Reproductive morphology Flowers Calyx
Sepals linear-lanceolate, 3-6 × 1-2 mm., deciduous.
Morphology Reproductive morphology Flowers Corolla
Petals lemon-yellow to orange-yellow, 4-8 × 3-6 mm.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 8-10 in 2 whorls; filaments up to 3 mm. long; anthers 0.5-1 mm. long.
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary globular, 1 mm. across; style very short or absent; stigma pyramidal, 0.8-1.2 mm. long.
Morphology Reproductive morphology Fruits
Fruit disc-shaped; stalk ± 1 cm. long; mericarp 4-5 × 2-3 mm., provided with 2 pairs of spines.
Habitat
Open and disturbed places, often on sandy soils, becoming a troublesome weed; 0-2300 m.
Distribution
K1 K2 K3 K4 K5 K6 K7 T1 T2 T3 T4 T5 T6 T7 tropical and temperate regions throughout the world

Native to:

Afghanistan, Albania, Algeria, Altay, Angola, Assam, Austria, Baleares, Bangladesh, Benin, Botswana, Bulgaria, Burkina, Burundi, Buryatiya, Cambodia, Cameroon, Canary Is., Cape Provinces, Cape Verde, Caprivi Strip, Central African Repu, Central European Rus, Chad, China North-Central, China South-Central, China Southeast, Corse, Cyprus, Czechoslovakia, Djibouti, East Aegean Is., East Himalaya, Egypt, Eritrea, Ethiopia, France, Free State, Gambia, Ghana, Greece, Guinea, Gulf States, Hainan, Hungary, India, Inner Mongolia, Iran, Iraq, Italy, Ivory Coast, Japan, Kazakhstan, Kenya, Kirgizstan, Korea, Kriti, Krym, Kuwait, KwaZulu-Natal, Lebanon-Syria, Libya, Malawi, Maldives, Mali, Manchuria, Mauritania, Mongolia, Morocco, Mozambique, Mozambique Channel I, Myanmar, Namibia, Nepal, Niger, Nigeria, North Caucasus, Northern Provinces, Oman, Pakistan, Palestine, Portugal, Qinghai, Romania, Rwanda, Sardegna, Saudi Arabia, Senegal, Sicilia, Sinai, Socotra, Somalia, South China Sea, South European Russi, Spain, Sri Lanka, Sudan, Swaziland, Tadzhikistan, Taiwan, Tanzania, Thailand, Tibet, Togo, Transcaucasus, Tunisia, Turkey, Turkey-in-Europe, Turkmenistan, Tuva, Uganda, Ukraine, Uzbekistan, Vietnam, West Himalaya, West Siberia, Xinjiang, Yemen, Yugoslavia, Zambia, Zaïre, Zimbabwe

Introduced into:

Alabama, Argentina Northeast, Argentina Northwest, Argentina South, Arizona, Arkansas, Bolivia, British Columbia, California, Chile North, Colorado, Delaware, Ecuador, Florida, Galápagos, Georgia, Hawaii, Idaho, Illinois, Indiana, Iowa, Kansas, Kentucky, Louisiana, Maryland, Massachusetts, Mexico Northwest, Michigan, Minnesota, Mississippi, Missouri, Montana, Nebraska, Nevada, New Jersey, New Mexico, New York, North Carolina, North Dakota, Ohio, Oklahoma, Ontario, Oregon, Panamá, Pennsylvania, Peru, Primorye, South Carolina, South Dakota, St.Helena, Tennessee, Texas, Turks-Caicos Is., Uruguay, Utah, Washington, Wisconsin, Wyoming

English
(Land) calthrop, Burnut, Devil's thorn, Puncture vine

Tribulus terrestris L. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Oct 16, 2008 Crawford, F. [121], Namibia K000450349
Tanner, R. [517], Tanzania 16011.000
Headley, P.M. [196], Ethiopia 6049.067
Sieber [s.n.], Senegal K000417026
Gillies, L. [s.n.], Senegal K000417027
Heudelot, D., Senegal K000417028
Ecklon [751], South Africa K000417017
South Africa K000417019
Spruce, R. [5306], Ecuador K000201327
Spruce, R. [5953], Ecuador K000201328
Mueller, F J.H.Von. [s.n.], Australia K000725218
Nascimento, A. [246], Brazil K001138324
Hatschbach, G. [61960], Brazil K001138326
Mueller, F.J.H.von. [s.n.], Australia K000725219
Cooper, T. [2148], South Africa K000417018
Mueller, F J.H.Von. [s.n.], Australia K000725214
Chevalier, A.J.B. [1287], Mali K001089859
Spruce, R. [5953], Ecuador K000201326
Queiroz, L.P. [2605], Brazil K001138325
Spruce, R. [6406], Ecuador K000201330
Mueller, F J.H.Von. [s.n.], Australia K000725216
Mueller, F J.H.Von. [s.n.], Australia K000725217
Pickersgill, B. [72-55], Brazil K001138323
s.coll. [43], South Australia K000725215
Wallich, N. [Cat. no. 1191], Myanmar Tribulus lanuginosus K001112360
s.coll. [Cat. no. 6854] Tribulus lanuginosus K001124989
s.coll. [Cat. no. 6854] Tribulus lanuginosus K001124990
s.coll. [Cat. no. 6854] Tribulus lanuginosus K001124991
s.coll. [Cat. no. 1191] Tribulus lanuginosus K001112359
Wallich, N. [Cat. no. 1191], India Tribulus lanuginosus K001112358
s.coll. [Cat. no. 6854], India Tribulus lanuginosus K001124988
Kerner, A. [39], Hungary Tribulus orientalis K000725201 syntype

First published in Sp. Pl.: 387 (1753)

Accepted by

  • Abdulina, S.A. (1999). Spisok Sosudistykn Rastenii Kazakhstana: 1-187. Academy of Sciences, Almaty, Kazakhstan.
  • Ackerfield, J. (2015). Flora of Colorado: 1-818. BRIT Press.
  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Allred, K.W. (2012). Flora Neomexicana, ed. 2, 1: 1-599. Range Science Herbarium, Las Cruces, New Mexico.
  • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
  • Bailey, C. & al. (2015). Guide to the Vascular Plants of Tennessee: 1-813. University of Tennessee press.
  • Balkrishna, A. (2018). Flora of Morni Hills (Research & Possibilities): 1-581. Divya Yoga Mandir Trust.
  • Barooah, C. & Ahmed, I. (2014). Plant diversity of Assam. A checklist of Angiosperms and Gymnosperms: 1-599. Assam science technology and environment council, India.
  • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
  • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
  • Boulos, L. (2000). Flora of Egypt 2: 1-352. Al Hadara Publishing, Cairo.
  • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
  • Brako, L. & Zarucchi, J.L. (1993). Catalogue of the Flowering Plants and Gymnosperms of Peru Monographs in Systematic Botany from the Missouri Botanical Garden 45: i-xl, 1-1286. Missouri Botanical Garden.
  • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
  • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
  • Cossu, T.A, Camarda, I. & Brundu, G. (2014). A catalogue of non-native weeds in irrigated crops in Sardinia (Italy) Webbia; Raccolta de Scritti Botanici 69: 145-156.
  • Danin, A. & Fragman- Sapir, O. (2019). Flora of Israel Online http://flora.org.il/en/plants/.
  • Daoud, H.S. in Al-Rawi, A. (1985). Flora of Kuwait 1: 1-224. Alden Press Ltd., U.K.
  • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
  • Dassanayake (ed.) (1988). A Revised Handbook to the Flora of Ceylon 6: 1-424. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
  • Davis, P.H. (ed.) (1967). Flora of Turkey and the East Aegean Islands 2: 1-581. Edinburgh University Press, Edinburgh.
  • Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A., & Tzanoudakis, D. (2013). Vascular plants of Greece. An annotated checklist: 1-372. Botanic gardens and botanical museum Berlin-Dahlem, Berlin and Hellenic botanical society, Athens.
  • Dy Phon, P. (2000). Dictionnaire des plantes utilisées au Cambodge: 1-915. chez l'auteur, Phnom Penh, Cambodia.
  • Edwards, S., Tadesse, M., Demissew, S. & Hedberg, I. (eds.) (2000). Flora of Ethiopia and Eritrea 2(1): 1-532. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • El Hadidi, N. (1985). Flora of Tropical East Africa, Zygophyllaceae: 1-14.
  • Exell, A.W., Fernandes, A. & Wild, H. (eds.) (1963). Flora Zambesiaca 2(1): 1-351. Royal Botanic Gardens, Kew.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Garcillán, P.P. & al. (2013). Plantas no nativas naturalizadas de la península de Baja California, México Botanical Sciences 91: 461-475.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Ghazanfar, S.A. (1992). An Annotated Catalogue of the Vascular Plants of Oman and their Vernacular names Scripta Botanica Belgica 2: 1-153.
  • Grierson, A.J.C. & Long, D.G. (1987). Flora of Bhutan 1(3): 466-834. Royal Botanic Gardens, Edinburgh.
  • Grubov, V.I. (2001). Key to the vascular plants of Mongolia (with an atlas) 2: 1-817. Academy of Sciences, Mongolian People's Republic, Ulaan Bator.
  • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
  • Hara, H., Stearn, W.T. & Williams, H.J. (1979). An Enumeration of the Flowering Plants of Nepal 2: 1-220. Trustees of British Museum, London.
  • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
  • Iwatsuki, K., Boufford, D.E. & Ohba, H. (eds.) (2001). Flora of Japan IIb: 1-550. Kodansha Ltd., Tokyo.
  • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
  • Jones, R.L. (2005). Plant life of Kentucky. An illustrated guide to the vascular flora: 1-833. The universitry press of Kentucky.
  • Jongbloed, M., Western, R.A. & Boer, B. (2000). Annotated Check-list for plants in the U.A.E.: 1-90. Zodiac Publishing, Dubai.
  • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181. Missouri Botanical Garden.
  • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
  • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
  • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
  • Lambdon, P. (2012). Flowering plants & ferns of St Helena: 1-624. Pisces publications for St Helena nature conservation group.
  • Larson, J., Reif, B., Nelson, B.E. & Hartman, R.L. (2014). Floristic studies in North Central New Mexico, U.S.A. the Sange de Cristo mountains Journal of the Botanical Research Institute of Texas 8: 271-303.
  • Lazkov, G.A. & Sultanova, B.A. (2011). Checklist of vascular plants of Kyrgyzstan Norrlinia 24: 1-166.
  • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
  • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
  • Litvinskaya, S.A. & Murtazaliev, R.A. (2013). Flora of the Northern Caucasus: An Atlas and Identification Book: 1-688. Fiton XXI.
  • Llewellyn, O.A., Hall, M., Miller, A.G., Al-Abbasi, T.M., Al-Wetaid, A.H., Al-Harbi, R.J. & Al-Shammari, K.F. (2011). Important plant areas in the Arabian peninsula: 4. Jabal Aja Edinburgh Journal of Botany 68: 199-224.
  • Malyschev, L.I., Peshkova, G.A. & Baikov, K.S. (eds.) (2007). Flora of Siberia 14: 1-210. Scientific Publishers, Inc., Enfield, Plymouth.
  • Meena, S.L. (2012). A checklist of the vascular plants of Banaskantha district, Gujarat, India Nelumbo 54: 39-91.
  • Meikle, R.D. (1977). Flora of Cyprus 1: 1-832. The Bentham-Moxon Trust Royal Botanic Gardens, Kew.
  • Memon, R.A. & al. (2014). Illustrated weed flora of Cotton crop of Khairpur district, Sindh, Pakistan Pakistn Journal of Botany 46: 5-12.
  • Miller, A.G. & Morris, M. (2004). Ethnoflora of Soqotra Archipelago: 1-759. The Royal Botanic Garden, Edinburgh.
  • Mohlenbrock, R.H. (2014). Vascular Flora of Illinois. A Field Guide, ed. 4: 1-536. Southern Illinois University Press, Carbondale.
  • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
  • Mosti, S., Raffaelli, M. & Tardelli, M. (2012). Contributions to the flora of central-southern Dhofar (Sultanate of Oman) Webbia; Raccolta de Scritti Botanici 67: 65-91.
  • Musselman, L.J. (2011). Checklist of Plants of Lebanon and Syria http://ww2.odu.edu/~lmusselm/plant/lebsyria/Checklist%20of%20Lebanon%20Plants.pdf.
  • Nasir, E. & Ali, S.I. (eds.) (1970-1995). Flora of West Pakistan 1-131.
  • Onana, J.M. (2011). The vascular plants of Cameroon a taxonomic checklist with IUCN assessments: 1-195. National herbarium of Cameroon, Yaoundé.
  • Ovchinnikov, P.N. (ed.) (1981). Flora Tadzhikskoi SSR 6: 1-725. Izd-vo Akademii nauk SSSR, Moskva.
  • Peschkova, G.A. (ed.) (2006). Flora of Siberia 10: 1-314. Scientific Publishers, Inc., Enfield, Plymouth.
  • Prakash, L. & Balasubramanian, P. (2018). Invasive alien flora of Sathyamangalam Tiger Reserve in Southern Eastern Ghats, India Indian Forester 144: 857-862.
  • Probatova, N.S. & al. (2006). Flora Rossiiskogo Dal'nego Vostoka. Dopolneniya i izmeneniya k izdaniyu Sosudistye rasteniya sovetskogo Dal'nego Vostoka t. 1-8 (1985-1996): 1-454. Russian academy of sciences Far Eastern branch.
  • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.
  • Santisuk, T. & Larsen, K. (eds.) (2000). Flora of Thailand 7(2): 253-349. The Forest Herbarium, Royal Forest Department.
  • Shaheen, H., Qureshi, R., Akram, A., Gulfraz, M. & Potter, D. (2014). A preliminary floristic checklist of Thal desert Punjab, Pakistan Pakistn Journal of Botany 46: 13-18.
  • Shishkin, B.K. (ed.) (1950). Flora Turkmenii 5: 1-271. Turkmenskoe gosudarstvennoe izd., Ashkhabad.
  • Sikarwar, R.L.S. (2014). Angiosperm diversity assessment of Chitrakootthe legendary place of Vindhyan range, India Journal of Economic and Taxonomic Botany 38: 563-619.
  • Smith, A.C. (1985). Flora Vitiensis Nova. A new flora for Fiji (Spermatophytes only) 3: 1-758. Pacific Tropical Botanical Garden, Lawai.
  • Townsend, C.C. & Guest, E. (eds.) (1980). Flora of Iraq 4(1): 1-628. Ministry of Agriculture & Agrarian Reform, Baghdad.
  • Troupin, G. (ed.) (1983). Flora du Rwanda 2: 1-603. Musee Royal de l'Afrique Centrale.
  • Tutin, T.G. & al. (eds.) (1968). Flora Europaea 2: 1-469. Cambridge University Press.
  • Vvedensky, A.I. (ed.) (1959). Flora Uzbekistana 4: 1-507. Izd-va Akademii nauk Uzbekskoi SSR, Tashkent.
  • Werier, D. (2017). Catalogue of the Vascular plants of New York state Memoirs of the Torrey Botanical Club 27: 1-542.
  • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
  • Wu, Z. & Raven, P.H. (eds.) (2008). Flora of China 11: 1-622. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
  • Zervous, S., Raus, T. & Yannitsaros, A. (2009). Additons to the flora of the island of Kalimnos (SE Aegean, Greece) Willdenowia 39: 165-177.

Literature

Flora of West Tropical Africa

  • Chev. Bot. 96
  • Engl. in E. & P. Pflanzenfam. 19A: 176 (1931).
  • —F.T.A. 1: 283

Kew Backbone Distributions

  • Abdulina, S.A. (1999). Spisok Sosudistykn Rastenii Kazakhstana: 1-187. Academy of Sciences, Almaty, Kazakhstan.
  • Ackerfield, J. (2015). Flora of Colorado: 1-818. BRIT Press.
  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Aké Assi, L. (2002). Flore de la Côte-d'Ivoire: catalogue systématique, biogéographie et écologie. II Boissiera 58: 1-401.
  • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
  • Bailey, C. & al. (2015). Guide to the Vascular Plants of Tennessee: 1-813. University of Tennessee press.
  • Barooah, C. & Ahmed, I. (2014). Plant diversity of Assam. A checklist of Angiosperms and Gymnosperms: 1-599. Assam science technology and environment council, India.
  • Barry, J. P. & Celles, J.S. (1991). Flore de Mauritanie 1: 1-359. Centre Regional de Documentation Pedagogique, Nice.
  • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
  • Boulos, L. (2000). Flora of Egypt 2: 1-352. Al Hadara Publishing, Cairo.
  • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTROM, Bangui.
  • Chang, C.S., Kim, H. & Chang, K.S. (2014). Provisional checklist of vascular plants for the Korea peninsula flora (KPF): 1-660. DESIGNPOST.
  • Correa A., Mireya D. Galdames, Carmen Correa A., M. D., C. Galdames & M. S. de Stapf (2004). Catálogo de las Plantas Vasculares de Panamá: 1-599. Smithsonian Tropical Research Institute.
  • Correll, D.S. & Correll, H.B. (1982). Flora of the Bahama Archipelago: 1-1692. J.Cramer, Vaduz.
  • Danin, A. & Fragman- Sapir, O. (2019). Flora of Israel Online http://flora.org.il/en/plants/.
  • Daoud, H.S. in Al-Rawi, A. (1985). Flora of Kuwait 1: 1-224. Alden Press Ltd., U.K.
  • Dassanayake (ed.) (1988). A Revised Handbook to the Flora of Ceylon 6: 1-424. Oxford & IBH Publishing Co. PVT. LTD., New Delhi, Calcutta.
  • Davis, P.H. (ed.) (1967). Flora of Turkey and the East Aegean Islands 2: 1-581. Edinburgh University Press, Edinburgh.
  • Dobignard, A. & Chatelain, C. (2013). Index synonymique de la flore d'Afrique du nord 5: 1-451. Éditions des conservatoire et jardin botaniques, Genève.
  • Edwards, S., Tadesse, M., Demissew, S. & Hedberg, I. (eds.) (2000). Flora of Ethiopia and Eritrea 2(1): 1-532. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • El Hadidi, M.N. (1972). Flora Iranica 98: 1-32. Akademische Druck- u. Verlagsanstalt, Graz.
  • El Hadidi, N. (1985). Flora of Tropical East Africa, Zygophyllaceae: 1-14.
  • Exell, A.W., Fernandes, A. & Wild, H. (eds.) (1963). Flora Zambesiaca 2(1): 1-351. Royal Botanic Gardens, Kew.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Flora of North America Editorial Committee (2016). Flora of North America North of Mexico 12: 1-603. Oxford University Press, New York, Oxford.
  • Fosberg, F.R. (1957). The Maldive islands, Indian Ocean Atoll Research Bulletin 58: 1-37.
  • Garcillán, P.P. & al. (2013). Plantas no nativas naturalizadas de la península de Baja California, México Botanical Sciences 91: 461-475.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Grierson, A.J.C. & Long, D.G. (1987). Flora of Bhutan 1(3): 466-834. Royal Botanic Gardens, Edinburgh.
  • Grubov, V.I. (2001). Key to the vascular plants of Mongolia (with an atlas) 2: 1-817. Academy of Sciences, Mongolian People's Republic, Ulaan Bator.
  • Hansen, A. & Sunding, P. (1993). Flora of Macaronesia. Checklist of vascular plants. 4. revised edition Sommerfeltia 17: 1-295.
  • Hara, H., Stearn, W.T. & Williams, H.J. (1979). An Enumeration of the Flowering Plants of Nepal 2: 1-220. Trustees of British Museum, London.
  • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
  • Iwatsuki, K., Boufford, D.E. & Ohba, H. (eds.) (2001). Flora of Japan IIb: 1-550. Kodansha Ltd., Tokyo.
  • Jaramillo Díaz, P. & Guézou, A. (2017). CDF Checklist of Galapagos Vascular Plants - FCD Lista de especies de Plantas Vasculares de Galápagos http://www.darwinfoundation.org/datazone/checklists/vascular-plants/.
  • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
  • Jones, R.L. (2005). Plant life of Kentucky. An illustrated guide to the vascular flora: 1-833. The universitry press of Kentucky.
  • Jongbloed, M., Western, R.A. & Boer, B. (2000). Annotated Check-list for plants in the U.A.E.: 1-90. Zodiac Publishing, Dubai.
  • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181. Missouri Botanical Garden.
  • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
  • Kral, R., Diamond, A.R., Ginzbarg, S.L., Hansen, C.J., Haynes, R.R., Keener, B.R., Lelong, M.G., Spaulding, D.D. & Woods, M. (2011). Annotated checklist of the vascular plants of Alabama: 1-112. Botanical reseach institute of Texas.
  • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590. Smithsonian Institution.
  • Lambdon, P. (2012). Flowering plants & ferns of St Helena: 1-624. Pisces publications for St Helena nature conservation group.
  • Larson, J., Reif, B., Nelson, B.E. & Hartman, R.L. (2014). Floristic studies in North Central New Mexico, U.S.A. the Sange de Cristo mountains Journal of the Botanical Research Institute of Texas 8: 271-303.
  • Lazkov, G.A. & Sultanova, B.A. (2011). Checklist of vascular plants of Kyrgyzstan Norrlinia 24: 1-166.
  • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Lewalle, J. (1970). Liste floristique et répartition altitudinale de la flore du Burundi occidental: 1-84. Université officielle de Bujumbura.
  • Lisowski, S. (2009). Flore (Angiospermes) de la République de Guinée Scripta Botanica Belgica 41: 1-517.
  • Litvinskaya, S.A. & Murtazaliev, R.A. (2013). Flora of the Northern Caucasus: An Atlas and Identification Book: 1-688. Fiton XXI.
  • Llewellyn, O.A., Hall, M., Miller, A.G., Al-Abbasi, T.M., Al-Wetaid, A.H., Al-Harbi, R.J. & Al-Shammari, K.F. (2011). Important plant areas in the Arabian peninsula: 4. Jabal Aja Edinburgh Journal of Botany 68: 199-224.
  • Lê, T.C. (2003). Danh l?c các loài th?c v?t Vi?t Nam 2: 1-1203. Hà N?i : Nhà xu?t b?n Nông nghi?p.
  • Malyschev, L.I., Peshkova, G.A. & Baikov, K.S. (eds.) (2007). Flora of Siberia 14: 1-210. Scientific Publishers, Inc., Enfield, Plymouth.
  • Meena, S.L. (2012). A checklist of the vascular plants of Banaskantha district, Gujarat, India Nelumbo 54: 39-91.
  • Meikle, R.D. (1977). Flora of Cyprus 1: 1-832. The Bentham-Moxon Trust Royal Botanic Gardens, Kew.
  • Memon, R.A. & al. (2014). Illustrated weed flora of Cotton crop of Khairpur district, Sindh, Pakistan Pakistn Journal of Botany 46: 5-12.
  • Miller, A.G. & Morris, M. (2004). Ethnoflora of Soqotra Archipelago: 1-759. The Royal Botanic Garden, Edinburgh.
  • Mohlenbrock, R.H. (2014). Vascular Flora of Illinois. A Field Guide, ed. 4: 1-536. Southern Illinois University Press, Carbondale.
  • Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
  • Mosti, S., Raffaelli, M. & Tardelli, M. (2012). Contributions to the flora of central-southern Dhofar (Sultanate of Oman) Webbia; Raccolta de Scritti Botanici 67: 65-91.
  • Musselman, L.J. (2011). Checklist of Plants of Lebanon and Syria http://ww2.odu.edu/~lmusselm/plant/lebsyria/Checklist%20of%20Lebanon%20Plants.pdf.
  • Nasir, E. & Ali, S.I. (eds.) (1970-1995). Flora of West Pakistan 1-131.
  • Onana, J.M. (2011). The vascular plants of Cameroon a taxonomic checklist with IUCN assessments: 1-195. National herbarium of Cameroon, Yaoundé.
  • Ovchinnikov, P.N. (ed.) (1981). Flora Tadzhikskoi SSR 6: 1-725. Izd-vo Akademii nauk SSSR, Moskva.
  • Peschkova, G.A. (ed.) (2006). Flora of Siberia 10: 1-314. Scientific Publishers, Inc., Enfield, Plymouth.
  • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Probatova, N.S. & al. (2006). Flora Rossiiskogo Dal'nego Vostoka. Dopolneniya i izmeneniya k izdaniyu Sosudistye rasteniya sovetskogo Dal'nego Vostoka t. 1-8 (1985-1996): 1-454. Russian academy of sciences Far Eastern branch.
  • Quézel, P. (1958). Mission Botanique au Tibesti: 1-357. Université d'Alger.
  • Robyns, W. & al. (eds.) (1948-1963). Flore du Congo Belge et du Ruanda-Urundi 1-10.
  • Santisuk, T. & Larsen, K. (eds.) (2000). Flora of Thailand 7(2): 253-349. The Forest Herbarium, Royal Forest Department.
  • Shishkin, B.K. (ed.) (1950). Flora Turkmenii 5: 1-271. Turkmenskoe gosudarstvennoe izd., Ashkhabad.
  • Thulin, M. (ed.) (1993). Flora of Somalia 1: 1-493. The Royal Botanic Gardens, Kew.
  • Troupin, G. (ed.) (1983). Flora du Rwanda 2: 1-603. Musee Royal de l'Afrique Centrale.
  • Turner, I.M., Xing, F. & Corlett, R.T. (2000). An annotated check-list of the vascular plants of the South China Sea and its shores The Raffles bulletin of Zoology, Suppl. 8: 23-116.
  • Tutin, T.G. & al. (eds.) (1968). Flora Europaea 2: 1-469. Cambridge University Press.
  • Vvedensky, A.I. (ed.) (1959). Flora Uzbekistana 4: 1-507. Izd-va Akademii nauk Uzbekskoi SSR, Tashkent.
  • Wagner, W.L., Herbst, D.R. & Sohmer, S.H. (1999). Manual of the Flowering Plants of Hawai'i, rev. ed., 2: 989-1918. University of Hawai'i Press, Bishop Museum Press.
  • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
  • Wu, Z. & Raven, P.H. (eds.) (2008). Flora of China 11: 1-622. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).
  • Zervous, S., Raus, T. & Yannitsaros, A. (2009). Additons to the flora of the island of Kalimnos (SE Aegean, Greece) Willdenowia 39: 165-177.
  • Zuloaga, F.O., Morrone, O. , Belgrano, M.J., Marticorena, C. & Marchesi, E. (eds.) (2008). Catálogo de las Plantas Vasculares del Cono Sur Monographs in Systematic Botany from the Missouri Botanical Garden 107: 1-3348. Missouri Botanical Garden.

Flora of Somalia

  • Flora Somalia, Vol 1, (1993) Author: by M. Thulin [updated by M. Thulin 2008]

Flora of Tropical East Africa

  • A. Schreiber in Prodr. Fl. SW.-Afr. 65: 6 (1966).
  • A.J. Jex-Blake, Some Wild Flowers of Kenya p. 21 (1948).
  • E. Lind & A. Tallantire, Some Common Flowering Plants of Uganda p. 63 (1962).
  • Engler in A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien, ed. 2, 19a: 176 (1931).
  • Engler in A. Engler & O. Drude, Die Vegetation Der Erde, IX, Pflanzenwelt Afrikas 3 (1): 736 (1915)
  • F. W. Andr., The Flowering Plants of the Anglo-Egyptian Sudan 1: 125 (1950).
  • G. Gilbert in Flore du Congo Belge et du Ruanda-Urundi, 7: 64 (1958).
  • H. Perrier in Fl. Madag. 103:4, fig. 1/1-3 (1952).
  • Hadidi in Tackholm, Students’ Fl. Egypt, ed. 2: 313, t. 100B (1974).
  • Hanid in A.D.Q. Agnew, Upland Kenya Wild Flowers p. 140 (1974).
  • Keay in Flora of West Tropical Africa, ed. 2, 1: 363 (1954).
  • L., Sp. Pl.: 387 (1753).
  • Launert in Flora Zambesiaca 2: 126, t. 19B (1963).
  • Letouzey in Fl. Cameroun 1: 155 (1963), pro parte
  • Letouzey in Fl. Gabon 6: 111 (1963), pro parte.
  • Oliv. in Flora of Tropical Africa 1: 283 (1868), pro parte.
  • Schweickerdt in Bothalia 3: 172 (1937).

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