1. Family: Zygophyllaceae R.Br.
    1. Genus: Tribulus L.
      1. Tribulus zeyheri Sond.

        This species is accepted, and its native range is Kenya to S. Africa.

    [FZ]

    Zygophyllaceae, E. Launert. Flora Zambesiaca 2:1. 1963

    Habit
    Perennial herb with spreading prostrate or somewhat ascending branches, finely hirsute with longer scattered bristle-like bulbous-based hairs on all vegetative parts; branches up to 1·2 m. long.
    Leaves
    Leaves unequal; the larger up to 9 cm. long, with up to 9 pairs of leaflets; the smaller up to 5 cm. long, with up to 4 pairs of leaflets.
    Leaflets
    Leaflets 4–20 × 2–11 mm., oblique, very variable, oblong, oblong-ovate or ovate, acute or subacute, pubescent on both surfaces or glabrous above, often ciliate.
    Stipules
    Stipules up to 10 mm. long, linear-lanceolate to obliquely ovate, ciliate.
    Peduncles
    Peduncle 1·5 to 2 times as long as the subtending leaf.
    Calyx
    Sepals 5–10 (12) × 1·8–2 mm., linear-lanceolate, acute, densely pubescent.
    Corolla
    Petals bright yellow, 10–20 (25) mm. long, up to 2·5 times the length of the sepals, broadly cuneate.
    Stamens
    Filaments 3–4 mm. long, anthers (1·2) 1·5–2·5 (3) mm. long.
    Pistil
    Style short; stigma 2–3 mm. long, slender, pyramidal.
    Ovary
    Ovary with stiff bristle-like tubercule-based hairs.
    Glands
    Intrastaminal glands connate, forming a shallow cup at the base of the ovary.
    Fruits
    Fruit breaking up into 5 cocci, each usually with 4 (sometimes 6) equal spines, or spines reduced to thick warts, tuberculate on the dorsal crest and often laterally compressed.
    [FTEA]

    Zygophyllaceae, M. Nabil El Hadidi (Faculty of Science, Cairo University). Flora of Tropical East Africa. 1985

    Habit
    Annual or biennial hirsute herb, with all the vegetative parts covered by a fine indumentum and bristle-like bulbous-based hairs; branches prostrate, up to 60 cm. long; internodes cylindrical, 2.5-8 cm. long and 2-3 mm. in diameter.
    Leaves
    Leaves 2.5-8 cm. long; leaflets 5-6 pairs, ovate, 7-10 × 4-5 mm.; stipules ovate-lanceolate, 3-5 × 1-2 mm., soon deciduous.
    Flowers
    Flowers 15-50 mm. across; pedicel 15-40 mm., 1.5-2 times as long as the subtending leaf.
    Calyx
    Sepals linear-lanceolate, 6-12 × 1-3 mm., densely hairy, deciduous.
    Corolla
    Petals yellow, 7-22 × 5-18 mm.
    Stamens
    Stamens 8-10 in 2 whorls; filaments 2-6 mm. long; anthers up to 2 mm. long.
    Pistil
    Ovary globular to pyriform; style short, 0.5-1.5 mm. long; stigma lanceolate, 1.5-3 mm. long.
    Fruits
    Fruit disc-shaped; stalk 1.5-5 cm. long; mericarp 4-10 × 2-6 mm., provided with 2 pairs of spines and sometimes a number of spinules.

    Images

    Distribution

    Native to:

    Angola, Botswana, Cape Provinces, Free State, Kenya, Namibia, Northern Provinces, Tanzania, Uganda, Zimbabwe

    Tribulus zeyheri Sond. appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Zeyher [272], South Africa K000417022 Unknown type material
    Zeyher [272], South Africa K000417023 Unknown type material

    First published in W.H.Harvey & auct. suc. (eds.), Fl. Cap. 1: 353 (1860)

    Accepted by

    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • El Hadidi, N. (1985). Flora of Tropical East Africa, Zygophyllaceae: 1-14.
    • Exell, A.W., Fernandes, A. & Wild, H. (eds.) (1963). Flora Zambesiaca 2(1): 1-351. Royal Botanic Gardens, Kew.

    Literature

    Kew Backbone Distributions
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
    • El Hadidi, N. (1985). Flora of Tropical East Africa, Zygophyllaceae: 1-14.
    • Exell, A.W., Fernandes, A. & Wild, H. (eds.) (1963). Flora Zambesiaca 2(1): 1-351. Royal Botanic Gardens, Kew.
    Flora of Tropical East Africa
    • A. Schreiber in Prod. Fl. SW.-Afr. 65: 6 (1966), pro parte.
    • Launert in Flora Zambesiaca 2: 128, t. 19A (1963), pro parte.
    • Excell & Mendonça in Consp. Fl. Angol. 1: 255 (1951), pro parte.
    • Schweick. in Bothalia 3: 168-169 (1937), pro parte.
    • Engler in A. Engler & K. Prantl, Die Natürlichen Pflanzenfamilien, ed. 2, 19a: 176 (1931).
    • Engler in A. Engler & O. Drude, Die Vegetation Der Erde, IX, Pflanzenwelt Afrikas 3 (1): 736 (1915)
    • Sond. in Fl. Cap. 1: 353 (1860).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0