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  1. Family: Rubiaceae Juss.
    1. Genus: Psydrax Gaertn.
      1. Psydrax subcordatus (DC.) Bridson

        This species is accepted, and its native range is Tropical Africa.

    [FZ]

    Rubiaceae, D. Bridson. Flora Zambesiaca 5:2. 1998

    Habit
    Tree 5–15 m tall, with a palm-like habit, associated with ants, the branches often hollow and swollen with access pores present; young branches distinctly square in cross section, often slightly winged on the angles, glabrous to pubescent.
    Leaves
    Leaf blades 9.5–22(30) × 4.5–16.5 cm, oblong to ovate or sometimes broadly ovate, shortly acuminate at apex, obtuse to rounded or truncate to subcordate at base, glabrous or sometimes sparsely pubescent on upper surface, glabrous or pubescent on nerves or sometimes sparsely pubescent beneath, papery to subcoriaceous; lateral nerves in 7–10 main pairs; tertiary nerves moderately conspicuous beneath; domatia present as hair-lined cavities in the axils of lateral and sometimes tertiary nerves; petioles 10–15 mm long, glabrous or pubescent; stipules 4–10 mm long (or 14–18 mm long on saplings or coppice growth), narrowly ovate to ovate-caducous.
    Flowers
    Flowers with a very unpleasant smell, 5-merous, borne in 80–120-flowered, dichotomous corymbose pedunculate cymes; true peduncle rather short, the portion above the bract being longer, 0.8–2(4) cm long altogether, pubescent or glabrous; pedicels 1.5–2.5 mm long, sparsely puberulous to pubescent; bracts 1–4 mm long, in connate pairs or sometimes free; bracteoles free, inconspicuous, or conspicuous and up to 3 mm long.
    Calyx
    Calyx tube c. 1 mm long, glabrous or sometimes pubescent towards the base; limb 0.5–1 mm long, truncate to dentate, ciliate, puberulous inside.
    Corolla
    Corolla white or creamy-yellow; tube 2–3.5 mm long, with a ring of deflexed hairs set above mid-point inside; lobes 2.5 × 1.25–1.5 mm, oblong ovate, acute.
    Anthers
    Anthers not persistent in mature flowers.
    Style
    Style 5–7 mm long.
    Pollen presenter
    Pollen presenter 1.5–2 mm long, ribbed; disk pubescent or sometimes glabrous.
    Fruits
    Fruit 6–8 × 9–12(14) mm, usually rather prominent at apex, black or greyish-black.
    Pyrene
    Pyrenes 6–8 × 4.5–6 × 3–4 mm, strongly curved, with a groove from point of attachment to the centre of the lateral face, with a shallow crest at the apex, cartilaginous, scarcely rugulose.
    [FTEA]

    Rubiaceae, B. Verdcourt. Flora of Tropical East Africa. 1976

    Habit
    Tree 5–15 m. tall, said to have a palm-like habit, associated with ants and often the branches hollow and swollen with access-pores present; young branches distinctly square, glabrous or pubescent.
    Leaves
    Leaf-blades oblong to ovate or sometimes broadly ovate, 9.5–22 cm. long, 4.5–16.5 cm. wide, shortly acuminate at apex, obtuse, rounded, truncate or sometimes cordate at base, glabrous or sometimes sparsely pubescent above, glabrous or pubescent on nerves or sparsely pubescent beneath, papery to subcoriaceous, dull to moderately shiny above; lateral nerves in 7–10 main pairs; tertiary nerves moderately conspicuous beneath; domatia present as hair-lined cavities in the axils of lateral and tertiary nerves; petioles 1–1.5 cm. long, glabrous or pubescent; stipules narrowly ovate to ovate, 0.4–1 cm. long (or 1.4 cm. on sapling or coppice growth), caducous.
    Flowers
    Flowers with extremely distasteful smell, 5-merous, borne in pedunculate, 80–120-flowered, dichotomous corymbose cymes; true peduncles rather short, portion above the bract usually much longer, 0.8–2(–4) cm. altogether, pubescent or glabrous; pedicels 1.5–2.5 mm. long, sparsely puberulous to pubescent; bracts connate or sometimes free, 1–4 mm. long; bracteoles free, conspicuous or inconspicuous, up to 3 mm. long.
    Calyx
    Calyx-tube ± 1 mm. long, glabrous or sometimes pubescent towards base; limb a dentate or truncate rim 0.5–1 mm. long, ciliate, puberulous inside.
    Corolla
    Corolla white or creamy yellow; tube 2–3.5 mm. long, with a ring of deflexed hairs set above mid-point inside; lobes oblong-ovate, 2.5 mm. long, 1.25–1.5 mm. wide, acute.
    Anthers
    Anthers not persistent in opened flower.
    Pistil
    Style 5–7 mm. long; stigmatic knob 1.5–2 mm. long, ribbed.
    Disc
    Disk pubescent or sometimes glabrous.
    Fruits
    Fruit black or greyish black, wider than long, 0.6–0.8 cm. long, 0.9–1.2 cm. wide, usually rather prominent at apex; pyrenes strongly curved, 6–8 mm. long, 4.5–6 mm. wide, 3–4 mm. thick, with a groove on either side from point of attachment to centre of lateral face and with a shallow crest around apex, cartilaginous, very slightly rugulose.
    [FWTA]

    Rubiaceae, F. N. Hepper & R. W. J. Keay. Flora of West Tropical Africa 2. 1963

    Habit
    A small or medium-size tree about 30 ft. or up to 80 ft. high with horizontal branches near the crown
    Slash
    Slash whitish
    Buttresses
    Buttresses at the base
    Flowers
    Flowers white, heavily scented
    Ecology
    In the more open parts of high forests.

    Images

    Distribution

    Native to:

    Angola, Benin, Burkina, Cabinda, Cameroon, Central African Repu, Chad, Congo, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Gulf of Guinea Is., Ivory Coast, Liberia, Nigeria, Rwanda, Senegal, Sierra Leone, Sudan, Togo, Uganda, Zambia, Zaïre

    Accepted Infraspecifics

    Synonyms

    Other Data

    Psydrax subcordatus (DC.) Bridson appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1984 Welwitsch [3149], Angola K000412577 syntype
    Jan 1, 1984 Welwitsch, Angola K000412576 syntype
    Troupin, G. [4766], Congo, DRC 847.000
    Gartlan, J.S. [17], Cameroon 7382.000
    Thomas, D.W. [743], Cameroon 41175.000
    Bridson, D. [337], Rwanda 41706.000

    Bibliography

    First published in Kew Bull. 40: 698 (1985)

    Accepted by

    • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo) Taxonomania. Revue de Taxonomie et de Nomenclature Botaniques 30: 1-307.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Figueiredo, E. (2007). The Rubiaceae of Cabinda (Angola) Botanical Journal of the Linnean Society 154: 455-495.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Kew Backbone Distributions

    • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Ruhsam, M in Ruhsam, M. (2005). Survey of all the Rubiaceae specimens at the Royal Botanic Gardens, Kew.
    • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
    • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.

    Flora of Tropical East Africa

    • Bridson in Fl. Rwanda 3: 150, fig. 44.3 (1985).
    • Bridson in Kew Bulletin 40: 698 (1985).

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0