1. Family: Santalaceae R.Br.
    1. Genus: Thesium L.
      1. Thesium fanshawei Hilliard

        This species is accepted, and its native range is Zambia.

    [FZ]

    Flora Zambesiaca. Vol. 9, Part 3. Polygonaceae-Myriaceae. Pope GV, Polhill RM, Martins ES. 2006.

    Type
    Type:Zambia, Machili, 14.vii.1960, Fanshawe 5784 (BR, holotype; K; LISC).
    Habit
    Perennial herb drying blackish, clearly with a woody caudex (only fragment seen) crowned with very short closely scaly shoots giving rise to several stems c. 300 mm long, 3.3 mm diameter at base, possibly ± prostrate (horizontal fide Fanshawe), simple below, soon fastigiately well branched, ultimate branches spreading ± at right angles to axes, each branch articulated to a small cushion, young parts weakly ribbed by decurrent leaves, all stems laxly leafy Perennial herb drying blackish, clearly with a woody caudex (only fragment seen) crowned with very short closely scaly shoots giving rise to several stems c. 300 mm long, 3.3 mm diameter at base, possibly ± prostrate (horizontal fide Fanshawe), simple below, soon fastigiately well branched, ultimate branches spreading ± at right angles to axes, each branch articulated to a small cushion, young parts weakly ribbed by decurrent leaves, all stems laxly leafy.
    Leaves
    Leaves at base of stem reduced to small scales, those subtending branches c. 6–10 × 1 mm, smaller on branchlets, spreading to ascending and often incurved, linear-subulate, slightly broadened at base, thick-textured, shallowly grooved on ventral surface at base, dorsal surface rounded, glabrous Leaves at base of stem reduced to small scales, those subtending branches c. 6–10 × 1 mm, smaller on branchlets, spreading to ascending and often incurved, linear-subulate, slightly broadened at base, thick-textured, shallowly grooved on ventral surface at base, dorsal surface rounded, glabrous.
    Flowers
    Flowers solitary, terminating short (c. 2–15 mm long) sparsely scaly spreading brachyblasts ± racemosely arranged, though the branchlet bearing brachyblasts may itself terminate in a flower Flowers solitary, terminating short (c. 2–15 mm long) sparsely scaly spreading brachyblasts ± racemosely arranged, though the branchlet bearing brachyblasts may itself terminate in a flower.
    Bracts
    Bracts 5 or 6 forming an involucre immediately below the flower, c. 1.3 × 0.7–0.8 mm, ovate-cuspidate, thick-textured, margins membranous, ciliolate Bracts 5 or 6 forming an involucre immediately below the flower, c. 1.3 × 0.7–0.8 mm, ovate-cuspidate, thick-textured, margins membranous, ciliolate.
    Style
    Style 1–1.4 mm long; stigma nearly reaching tip of hood
    Ecology
    Known only from type collections made in woodland.
    Note
    The specimens were distributed as T. fastigiatum, which differs in its ascending branches and long filiform closely scaly ultimate branchlets. The relationship of T. fanshawei appears to lie more with T. doloense Pilg. (from the environs of Kinshasha, Dem. Rep. Congo) and T. atrum. It may be distinguished from T. doloense by its branchlets spreading ± at right angles to the stems (not ascending at angle of 45°) smaller and differently shaped involucral bracts (c. 1.3 × 0.8 mm, ovate-cuspidate, versus 1.8–2 × 0.6–0.7 mm, lanceolate-cuspidate) and larger flowers with perianth tube 0.7–0.8 mm long (not c. 0.25 mm), anthers 0.4–0.7 mm long (not 0.25 mm) and style 1–1.4 mm (not 0.7 mm). It may be distinguished from T. atrum by the plant drying fuscous (not black), leaves subtending main branches linear-subulate, 6–10 × 1 mm (not all reduced to bracts, lanceolate to ovate-lanceolate, 1.2–4 × 0.8–1.2 mm).
    Distribution
    ZAM B Zambia
    Perianth
    Perianth “greeny-white”; tube 0.7–0.8 mm long, external glands rather obscure, ellipsoid; lobes 1–1.1 × 0.5–0.7 mm, triangular, subacute, hood c. 0.3 mm long, margins inflexed, somewhat erose in lower half. Perianth “greeny-white”; tube 0.7–0.8 mm long, external glands rather obscure, ellipsoid; lobes 1–1.1 × 0.5–0.7 mm, triangular, subacute, hood c. 0.3 mm long, margins inflexed, somewhat erose in lower half
    Stamens
    Stamens inserted at base of perianth lobes; filaments 0.3–0.4 mm long, hidden by anthers; anthers 0.4–0.7 mm long. Stamens inserted at base of perianth lobes; filaments 0.3–0.4 mm long, hidden by anthers; anthers 0.4–0.7 mm long
    Pistil
    Style 1–1.4 mm long; stigma nearly reaching tip of hood.
    Fruits
    Fruit (immature) c. 3 × 2 mm (but possibly galled), ribs and reticulations strongly raised. Fruit (immature) c. 3 × 2 mm (but possibly galled), ribs and reticulations strongly raised.

    Images

    Distribution

    Thesium fanshawei Hilliard appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1989 Fanshawe, D.B. [F5784], Zambia K000431516 isotype

    First published in Edinburgh J. Bot. 48: 53 (1991)

    Accepted by

    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • Edinb. J. Bot. 48: 53 (1991).
    Kew Backbone Distributions
    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0