1. Family: Santalaceae R.Br.
    1. Genus: Thesium L.
      1. Thesium inonoense Hilliard

        This species is accepted, and its native range is N. Zambia.

    [FZ]

    Flora Zambesiaca. Vol. 9, Part 3. Polygonaceae-Myriaceae. Pope GV, Polhill RM, Martins ES. 2006.

    Type
    Type:Zambia, Mbala Distr., Katwe road, Inono Stream, 18.v.1955, Richards 5758 (K, holotype; BR).
    Style
    Style 0.7–1.3 mm long; stigma reaching top of anthers
    Fruits
    Fruit 1.5 × 1.5 mm, buff-coloured, strongly ribbed and reticulate. Fruit 1.5 × 1.5 mm, buff-coloured, strongly ribbed and reticulate.
    Ecology
    “Steep bank above Inono Stream, yellow soil, among tall grass”.
    Note
    Mrs. Richards' collection was determined as Thesium bangweolense, but that species is a perennial herb with a stout caudex crowned with masses of very short closely scaly shoots from which arise many slender scaly stems (quite different from the solitary, remotely scaly stem of T. inonoense with its little tuft of roots), each flower is subtended by an involucre of bracts terminating a very short closely bracteate brachyblast, and the fruits are larger (c. 2.5 × 2 mm). The affinity of T. inonoense lies with T. fimbriatum, from which it differs in its smaller floral bracts, perianth and fruits, and sparsely (not densely) fringed perianth lobes.
    Distribution
    Zambia ZAM N Known only from the type collection.
    Habit
    Probably annual; stems c. 300–400 mm tall, c. 1.5 mm in diameter at base, solitary, simple below, virgately branched above, branches sharply ascending, strongly ribbed by raised vascular strands terminating in midrib of scale leaves and bracts, distantly leafy. Probably annual; stems c. 300–400 mm tall, c. 1.5 mm in diameter at base, solitary, simple below, virgately branched above, branches sharply ascending, strongly ribbed by raised vascular strands terminating in midrib of scale leaves and bracts, distantly leafy
    Leaves
    Leaves all reduced to scales, c. 0.8–1.2 × 0.4–0.6 mm, lanceolate, acuminate, closely appressed, midrib strongly raised on dorsal surface. Leaves all reduced to scales, c. 0.8–1.2 × 0.4–0.6 mm, lanceolate, acuminate, closely appressed, midrib strongly raised on dorsal surface
    Flowers
    Flowers sessile in axil of each bract, sometimes solitary, mostly up to 6 crowded on very short brachyblasts, arranged in long lax spiciform inflorescences forming a lax panicle. Flowers sessile in axil of each bract, sometimes solitary, mostly up to 6 crowded on very short brachyblasts, arranged in long lax spiciform inflorescences forming a lax panicle
    Bracts
    Bracts c. 1.5–1.8 × 1–1.25 mm, ovate-acuminate, margins membranous, minutely erose-ciliate, keeled by strongly raised midrib; bracteoles 2, similar to bracts. Bracts c. 1.5–1.8 × 1–1.25 mm, ovate-acuminate, margins membranous, minutely erose-ciliate, keeled by strongly raised midrib; bracteoles 2, similar to bracts
    Pedicel
    Pedicels wanting. Pedicels wanting
    Perianth
    Perianth “cream”, tube 0.3–0.6 mm long, external glands either wanting or represented by almost imperceptible points alternating with the perianth lobes; lobes 1.2–1.6 × 0.4 mm, subspathulate, apex subacute, hood c. 0.2 mm long, margins inflexed, margins sparsely fringed in lower half. Perianth “cream”, tube 0.3–0.6 mm long, external glands either wanting or represented by almost imperceptible points alternating with the perianth lobes; lobes 1.2–1.6 × 0.4 mm, subspathulate, apex subacute, hood c. 0.2 mm long, margins inflexed, margins sparsely fringed in lower half
    Stamens
    Stamens inserted at base of perianth lobes; filaments conspicuous, 0.5–0.8 mm long, adnate at base to perianth lobes; anthers 0.2–0.3 mm long. Stamens inserted at base of perianth lobes; filaments conspicuous, 0.5–0.8 mm long, adnate at base to perianth lobes; anthers 0.2–0.3 mm long
    Pistil
    Style 0.7–1.3 mm long; stigma reaching top of anthers.

    Images

    Distribution

    Thesium inonoense Hilliard appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1989 Richards, H.M. [5758], Zambia K000431519 holotype

    First published in Edinburgh J. Bot. 48: 54 (1991)

    Accepted by

    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • Edinb. J. Bot. 48: 54 (1991). TAB. 46, fig. F.
    Kew Backbone Distributions
    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0