1. Family: Santalaceae R.Br.
    1. Genus: Thesium L.
      1. Thesium singulare Hilliard

        This species is accepted, and its native range is Zambia.

    [FZ]

    Flora Zambesiaca. Vol. 9, Part 3. Polygonaceae-Myriaceae. Pope GV, Polhill RM, Martins ES. 2006.

    Type
    Type:Zambia, 40 km west of Kaoma (Mankoya), fr. 11.xi.1965, E.A. Robinson 6705 (LISC, holotype; K; SRGH).
    Bracts
    Bracts c. 1 × 0.6–0.8 mm, similar to the scale leaves; bracteoles 2, similar to bract and scarcely smaller Bracts c. 1 × 0.6–0.8 mm, similar to the scale leaves; bracteoles 2, similar to bract and scarcely smaller.
    Style
    Style c. 5 mm long; stigma nearly reaching top of perianth lobes
    Note
    Remarkable in habit and inflorescence: the stolons bear at short intervals a few crowded, very short, densely scaly vegetative buds similar to those developed on the caudex not only in this species but particularly in species nos. 19–26 below. These buds eventually develop into flowering stems. The stem of Thesium singulare branches ± virgately in its upper half, but each twig terminates in a solitary flower, stem growth continuing from the axil of a scale leaf immediately below the flower. The branching of the inflorescence axis is therefore monochasial. In the specimens available, up to 3 flowers are produced in each cyme, which terminates in a long sterile scaly shoot.
    Distribution
    ZAM B Zambia Known only from the type collection, made in “seasonally damp grassland” in Barotseland.
    Habit
    Perennial herb with a woody caudex c. 15 mm in diameter crowned with crowded, very short, scaly vegetative buds, also with stout woody stolons up to c. 4 mm in diameter, with more distant scaly buds, these eventually producing erect flowering stems, loosely tufted; stems 110–150 mm tall, very slender (less than 1.3 mm diameter), simple below, sparingly branched above, branches sharply ascending, longitudinally finely striate, ± distantly leafy, whole plant drying buff-coloured to greenish. Perennial herb with a woody caudex c. 15 mm in diameter crowned with crowded, very short, scaly vegetative buds, also with stout woody stolons up to c. 4 mm in diameter, with more distant scaly buds, these eventually producing erect flowering stems, loosely tufted; stems 110–150 mm tall, very slender (less than 1.3 mm diameter), simple below, sparingly branched above, branches sharply ascending, longitudinally finely striate, ± distantly leafy, whole plant drying buff-coloured to greenish
    Leaves
    Leaves all reduced to scales 1–2 × 0.8–1 mm, lanceolate, acuminate, midrib strongly raised on dorsal surface near base; scales at extreme base of stem closely imbricate, distant and closely appressed on expanded stem. Leaves all reduced to scales 1–2 × 0.8–1 mm, lanceolate, acuminate, midrib strongly raised on dorsal surface near base; scales at extreme base of stem closely imbricate, distant and closely appressed on expanded stem
    Flowers
    Flowers solitary, terminal but soon overtopped by a branch developing from the axil of a scale leaf immediately below the floral bract and so resembling a very lax 2–3-flowered spike, but technically a monochasial cyme, often terminating in a long very slender, sterile shoot. Flowers solitary, terminal but soon overtopped by a branch developing from the axil of a scale leaf immediately below the floral bract and so resembling a very lax 2–3-flowered spike, but technically a monochasial cyme, often terminating in a long very slender, sterile shoot
    Perianth
    Perianth “white”; tube 2.6–3 mm long, external glands conspicuous, ellipsoid, about half as long as tube, situated immediately above ovary; lobes 2.7–3 × 0.6–0.8 mm, oblong, ± obtuse, margins inflexed, weakly erose-fimbriate. Perianth “white”; tube 2.6–3 mm long, external glands conspicuous, ellipsoid, about half as long as tube, situated immediately above ovary; lobes 2.7–3 × 0.6–0.8 mm, oblong, ± obtuse, margins inflexed, weakly erose-fimbriate
    Stamens
    Stamens inserted near base of perianth lobes; filaments 0.9–1.3 mm long, conspicuous; anthers 0.8–0.9 mm long. Stamens inserted near base of perianth lobes; filaments 0.9–1.3 mm long, conspicuous; anthers 0.8–0.9 mm long
    Pistil
    Style c. 5 mm long; stigma nearly reaching top of perianth lobes.
    Fruits
    Fruit c. 2.5 × 2.5 mm, pallid, ribs strongly raised, reticulations coarse, strongly raised. Fruit c. 2.5 × 2.5 mm, pallid, ribs strongly raised, reticulations coarse, strongly raised.

    Images

    Distribution

    Thesium singulare Hilliard appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Jan 1, 1989 Robinson, E.A. [6705], Zambia K000431523 isotype

    First published in Edinburgh J. Bot. 48: 54 (1991)

    Accepted by

    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • Edinb. J. Bot. 48: 54 (1991). TAB. 46, fig. B.
    Kew Backbone Distributions
    • Pope, G.V., Polhill, R.M. & Martins, E.S. (eds.) (2006). Flora Zambesiaca 9(3): 1-277. Royal Botanic Gardens, Kew.

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0