Guadua Kunth

This genus is accepted, and its native range is Mexico to S. Tropical America, Trinidad.

[GB]

nonem

Morphology General Habit
Perennial. Rhizomes short (27), or elongated (1); pachymorph. Culms erect (18/26), or geniculately ascending (1/26), or arching (5/26), or leaning (2/26), or scandent (4/26); 300-1145-3000 cm long; woody; without nodal roots (1/20), or with aerial roots from the nodes (1/20), or with root thorns from the nodes (19/20). Culm-internodes terete (22/27), or semiterete (5/27). Culm-nodes without obvious supra-nodal ridge (2/7), or with distinct supra-nodal ridge (5/7). Lateral branches dendroid. Branch complement one (4/20), or two (8/20), or three (13/20), or several (10/20), or many (1/20); in a horizontal line (1/5), or in a clump (4/5); with 1 branch dominant (9/9); thinner than stem (11/11). Culm-sheaths persistent (2/12), or tardily deciduous (1/12), or deciduous but leaving a persistent girdle (1/12), or deciduous (9/12); without auricles (13/16), or auriculate (6/16). Culm-sheath blade inconspicuous (1/18), or lanceolate (4/18), or ovate (2/18), or triangular (12/18); as wide as sheath at base (2/2). Leaf-sheath auricles absent (21), or erect (2), or falcate (5). Ligule an eciliate membrane (21), or a ciliolate membrane (10). Leaf-blade base with a brief petiole-like connection to sheath. Leaf-blades persistent (25), or deciduous at the ligule (3); linear (9), or lanceolate (26), or elliptic (1), or oblong (3), or ovate (2). Leaf-blade midrib conspicuous (5/5). Leaf-blade venation without cross veins (20), or with obscure cross veins (1), or with distinct cross veins (8).
Morphology Reproductive morphology Inflorescences
Synflorescence simple (1/26), or bractiferous (25/26); scanty (9/25), or paniculate (1/25), or globose (1/25), or clustered at the nodes (15/25); in untidy tufts (14/14); with glumaceous subtending bracts (24/25), or spathaceous subtending bracts (1/25); with axillary buds at base of spikelet (20/20); leafy between clusters (5/15), or leafless between clusters (8/15), or leafless between branches (2/15). Inflorescence a panicle (1/1). Spikelets appressed (3/4), or ascending (4/4), or spreading (1/4), or deflexed (1/4). Fertile spikelets sessile (26/26).
Morphology Reproductive morphology Inflorescences Spikelets
Spikelets comprising 1 basal sterile florets (5/5), or 2 basal sterile florets (4/5); 1-6-17 fertile florets; with a barren rhachilla extension (1/26), or with diminished florets at the apex (26/26). Spikelets linear (16/26), or lanceolate (10/26), or elliptic (1/26), or oblong (3/26); laterally compressed (19/26), or subterete (7/26); 3-38.49-170 mm long; breaking up at maturity (26/26); disarticulating below each fertile floret (26/26). Rhachilla internodes glabrous (18/25), or pubescent (7/25).
Fertile
Spikelets comprising 1 basal sterile florets (5/5), or 2 basal sterile florets (4/5); 1-6-17 fertile florets; with a barren rhachilla extension (1/26), or with diminished florets at the apex (26/26). Spikelets linear (16/26), or lanceolate (10/26), or elliptic (1/26), or oblong (3/26); laterally compressed (19/26), or subterete (7/26); 3-38.49-170 mm long; breaking up at maturity (26/26); disarticulating below each fertile floret (26/26). Rhachilla internodes glabrous (18/25), or pubescent (7/25).
Morphology Reproductive morphology Inflorescences Bracts Glume
Glumes two (6/22), or several (16/22); persistent (21/22), or deciduous (1/22); shorter than spikelet (22/22). Lower glume ovate (7/7); 0.25-0.5167-0.8 length of upper glume; membranous (1/4), or coriaceous (3/4); without keels (3/3); 7-9 -veined (2/4), or 10 -veined (1/4), or 11 -veined (3/4), or 12-13 -veined (2/4), or 14-17 -veined (1/4). Lower glume surface smooth (21/22), or asperulous (1/22); glabrous (20/22), or pubescent (2/22). Lower glume apex obtuse (3/6), or acute (4/6), or rostrate (1/6); muticous (6/7), or mucronate (1/7). Upper glume lanceolate (2/17), or ovate (16/17); 0.5-0.65-0.8 length of adjacent fertile lemma; coriaceous (11/11); without keels (12/12); 7-9 -veined (2/9), or 10 -veined (1/9), or 11-12 -veined (6/9), or 13 -veined (7/9), or 14-15 -veined (6/9), or 16-17 -veined (5/9), or 18-19 -veined (1/9). Upper glume lateral veins with cross-veins (4/4). Upper glume surface smooth (21/22), or asperulous (1/22); glabrous (19/22), or puberulous (3/22), or pubescent (3/22). Upper glume apex obtuse (2/13), or acute (11/13), or rostrate (1/13); muticous (12/17), or mucronate (5/17).
Morphology Reproductive morphology Flowers Florets
Basal sterile florets barren (5/5); with palea (4/5), or without significant palea (2/5). Lemma of lower sterile floret lanceolate (2/5), or elliptic (1/5), or ovate (3/5); chartaceous (2/2); 10-14 -veined (1/4), or 15-16 -veined (2/4), or 17-20 -veined (1/4), or 23-27 -veined (1/4); obtuse (1/4), or acuminate (3/4); mucronate (5/5). Fertile lemma lanceolate (5/25), or elliptic (1/25), or ovate (22/25); coriaceous (26/26); without keel (26/26); 7-17-30 -veined. Lemma lateral veins obscure (1/5), or prominent (4/5). Lemma surface smooth (25/26), or scaberulous (1/26); glabrous (17/26), or puberulous (6/26), or pubescent (6/26), or pilose (2/26). Lemma margins flat (25/26), or convolute (1/26); exposing palea (25/26), or covering most of palea (1/26); eciliate (20/26), or ciliolate (3/26), or ciliate (3/26). Lemma apex obtuse (2/25), or acute (17/25), or acuminate (3/25), or apiculate (3/25); muticous (14/26), or mucronate (12/26), or awned (2/26); 1 -awned (1/1). Palea 0.75-0.9729-1 length of lemma; 6 -veined (1/9), or 8 -veined (2/9), or 9 -veined (6/9), or 10-11 -veined (5/9), or 12 -veined (3/9), or 13 -veined (2/9). Palea keels winged (26/26); eciliate (16/26), or pubescent (1/26), or ciliolate (6/26), or ciliate (3/26). Palea surface glabrous (20/26), or puberulous (1/26), or pubescent (5/26). Apical sterile florets resembling fertile though underdeveloped (23/26), or distinct from fertile (3/26); 1 in number (3/3); barren (1/1); rudimentary (2/3), or linear (1/3).
Morphology Reproductive morphology Flowers
Lodicules 3 (26/26); membranous (3/3); glabrous (19/26), or ciliate (7/26). Anthers 3 (2/26), or 6 (24/26). Stigmas 3 (26/26). Ovary unappendaged (19/26), or umbonate (7/26); pubescent on apex (2/3), or pubescent all over (1/3).
Morphology Reproductive morphology Fruits
Caryopsis with adherent pericarp (25/26), or fleshy pericarp (1/26); fusiform (4/7), or oblong (2/7), or ovoid (1/7), or obovoid (1/7). Embryo 0.2-0.2875-0.5 length of caryopsis. Hilum linear (1/1); 1 length of caryopsis.
Distribution
North America (6), or South America (27).

[GB]

nonem

Morphology General Habit
Perennial. Rhizomes absent (1), or short (138), or elongated (2); pachymorph (140/140). Culms erect (114/124), or geniculately ascending (3/124), or arching (2/124), or leaning (1/124), or scandent (4/124); 76-1047-3000 cm long; woody; without nodal roots (73/96), or with aerial roots from the nodes (11/96), or with root thorns from the nodes (26/96). Culm-internodes terete. Culm-nodes flush with internodes (21/31), or swollen (10/31); without obvious supra-nodal ridge (2/2). Lateral branches dendroid. Branch complement one (7/104), or two (6/104), or three (17/104), or several (76/104), or many (21/104); in a clump (19/19); with subequal branches (5/46), or 1 branch dominant (38/46), or 3 branches dominant (3/46); thinner than stem (17/17). Culm-sheaths persistent (4/99), or tardily deciduous (5/99), or deciduous but leaving a persistent girdle (2/99), or deciduous (89/99); without auricles (17/118), or auriculate (102/118). Culm-sheath blade inconspicuous (1/123), or lanceolate (25/123), or narrowly ovate (18/123), or ovate (16/123), or triangular (80/123); constricted at base (10/43), or narrower than sheath (16/43), or as wide as sheath at base (7/43), or cordate (10/43). Leaf-sheath auricles absent (81), or erect (10), or falcate (58). Ligule an eciliate membrane (121), or a ciliolate membrane (18), or a ciliate membrane (6). Leaf-blade base without a false petiole (1), or with a brief petiole-like connection to sheath (140). Leaf-blades persistent (132), or deciduous at the ligule (12); filiform (1), or linear (19), or lanceolate (126), or elliptic (1), or oblong (21), or ovate (4). Leaf-blade midrib evident (3/22), or conspicuous (19/22). Leaf-blade venation without cross veins (129), or with obscure cross veins (8), or with distinct cross veins (6). Leaf-blade margins cartilaginous (1/1).
Morphology Reproductive morphology Inflorescences
Synflorescence bractiferous (79/79); paniculate (1/79), or clustered at the nodes (78/79); in globose clusters (2/78), or stellate clusters (1/78), or oblong clusters (4/78), or untidy tufts (71/78); with glumaceous subtending bracts (59/75), or spathaceous subtending bracts (16/75); with axillary buds at base of spikelet (79/79); leafy between clusters (6/43), or leafless between clusters (36/43), or leafless between branches (3/43). Fertile spikelets sessile (2/2).
Morphology Reproductive morphology Inflorescences Spikelets
Spikelets comprising 1-6-13 fertile florets; without rhachilla extension (3/79), or with a barren rhachilla extension (2/79), or with diminished florets at the apex (74/79). Spikelets linear (17/79), or lanceolate (49/79), or elliptic (1/79), or oblong (4/79), or ovate (15/79); laterally compressed (59/79), or subterete (20/79); 3.32-24.63-75 mm long; breaking up at maturity (79/79); rhachilla deciduous (78/79), or rhachilla persistent (1/79); disarticulating below each fertile floret (77/78), or above glumes but not between florets (1/78); shedding paleas (1/1). Rhachilla internodes glabrous (59/76), or sparsely hairy (2/76), or pubescent (10/76), or pilose (6/76). Floret callus glabrous (1/1).
Fertile
Spikelets comprising 1-6-13 fertile florets; without rhachilla extension (3/79), or with a barren rhachilla extension (2/79), or with diminished florets at the apex (74/79). Spikelets linear (17/79), or lanceolate (49/79), or elliptic (1/79), or oblong (4/79), or ovate (15/79); laterally compressed (59/79), or subterete (20/79); 3.32-24.63-75 mm long; breaking up at maturity (79/79); rhachilla deciduous (78/79), or rhachilla persistent (1/79); disarticulating below each fertile floret (77/78), or above glumes but not between florets (1/78); shedding paleas (1/1). Rhachilla internodes glabrous (59/76), or sparsely hairy (2/76), or pubescent (10/76), or pilose (6/76). Floret callus glabrous (1/1).
Morphology Reproductive morphology Inflorescences Bracts Glume
Glumes both absent or obscure (1/78), or one the lower absent or obscure (8/78), or two (31/78), or several (45/78); persistent (43/43); shorter than spikelet (42/42). Lower glume oblong (1/22), or ovate (21/22); 0.7-0.75-0.8 length of upper glume; chartaceous (1/4), or coriaceous (3/4); without keels (5/5); 7 -veined (1/6), or 9 -veined (3/6), or 10-12 -veined (2/6), or 13 -veined (1/6), or 15-17 -veined (1/6). Lower glume surface glabrous (71/73), or pubescent (2/73). Lower glume apex obtuse (2/15), or acute (12/15), or acuminate (1/15); muticous (18/22), or mucronate (4/22). Upper glume lanceolate (2/34), or ovate (32/34); without auricles (77/78), or auriculate at base (1/78); 0.5 length of adjacent fertile lemma; chartaceous (1/2), or coriaceous (1/2); without keels (3/3), or 1-keeled (1/3); 7 -veined (1/5), or 9-10 -veined (2/5), or 11 -veined (3/5), or 12-13 -veined (2/5), or 22 -veined (1/5). Upper glume lateral veins with cross-veins (1/1). Upper glume surface smooth (77/78), or scabrous (1/78); glabrous (76/78), or pubescent (2/78). Upper glume apex obtuse (2/19), or acute (15/19), or acuminate (2/19); muticous (27/34), or mucronate (7/34).
Morphology Reproductive morphology Flowers Florets
Fertile lemma lanceolate (17/78), or elliptic (2/78), or oblong (2/78), or ovate (63/78); chartaceous (4/5), or coriaceous (1/5); without keel (77/79), or keeled (2/79); 7-17-32 -veined. Lemma lateral veins obscure (3/3), or distinct (1/3). Lemma surface smooth (77/79), or scabrous (2/79); glabrous (77/79), or puberulous (1/79), or pubescent (1/79). Lemma margins eciliate (57/79), or ciliolate (9/79), or ciliate (14/79). Lemma apex obtuse (7/74), or acute (56/74), or acuminate (8/74), or attenuate (1/74), or cuspidate (2/74), or apiculate (4/74); without ornament (76/79), or scabrous (1/79), or pubescent (2/79); muticous (62/79), or mucronate (16/79), or awned (1/79); 1 -awned (1/1). Palea 1-1.005-1.1 length of lemma; membranous (78/79), or chartaceous (1/79); 3-10-18 -veined; 2-keeled but the uppermost without keels (1/79), or 2-keeled (78/79). Palea keels separated (77/79), or approximate (1/79), or contiguous above a sulcus (1/79); wingless (78/79), or winged (1/79); smooth (77/79), or scabrous (2/79); eciliate (22/79), or pubescent (3/79), or ciliolate (24/79), or ciliate (31/79). Palea surface glabrous (68/79), or puberulous (2/79), or pubescent (4/79), or pilose (6/79), or hirsute (1/79). Palea apex entire (2/6), or dentate (5/6). Apical sterile florets resembling fertile though underdeveloped (68/74), or distinct from fertile (7/74); 1 in number (7/7); male (1/5), or barren (4/5); rudimentary (4/6), or linear (1/6), or elliptic (1/6).
Morphology Reproductive morphology Flowers
Lodicules absent (6/78), or 2 (3/78), or 3 (71/78); membranous (67/67); glabrous (13/73), or ciliate (60/73). Anthers 3 (1/79), or 6 (78/79), or 7 (1/79). Stigmas 1 (6/77), or 2 (13/77), or 3 (69/77); plumose (71/79), or sparsely hairy (3/79), or pubescent (6/79), or retrorsely barbellate (1/79). Ovary unappendaged (30/79), or umbonate (49/79); glabrous (7/54), or with a few apical hairs (3/54), or pubescent on apex (44/54), or pubescent all over (1/54).
Morphology Reproductive morphology Fruits
Caryopsis with adherent pericarp (52/52); fusiform (1/12), or ellipsoid (1/12), or oblong (5/12), or ovoid (4/12), or obovoid (1/12). Embryo 0.2 length of caryopsis.
Distribution
Africa (4), or Temperate Asia (86), or Tropical Asia (67), or Australasia (5), or Pacific (9), or North America (1), or South America (5).

Native to:

Argentina Northeast, Argentina Northwest, Belize, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Colombia, Costa Rica, Ecuador, El Salvador, French Guiana, Guatemala, Guyana, Honduras, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Southeast, Mexico Southwest, Nicaragua, Panamá, Paraguay, Peru, Suriname, Trinidad-Tobago, Uruguay, Venezuela

Introduced into:

Galápagos, Myanmar, Puerto Rico

Guadua Kunth appears in other Kew resources:

Date Reference Identified As Barcode Type Status Has image?
Jan 1, 1988 Nee, M. [34515], Brazil K001106364 Yes
Martinelli, G. [12008], Brazil K001106348 Yes
Martinelli, G. [12008], Brazil K001106349 Yes
Calderon, C.E. [2873], Brazil K001106366 Yes
Calderon, C.E. [2873], Brazil K001106367 Yes
Calderon, C.E. [2846], Brazil K001106409 Yes
Calderon, C.E. [2813], Brazil K001106411 Yes
Calderon, C.E. [2813], Brazil K001106413 Yes
Calderon, C.E. [2833], Brazil K001106416 Yes
Trail, J.W.H. [1281], Brazil K001106399 Yes
Calderon, C.E. [2833], Brazil K001106414 Yes
Calderon, C.E. [2910], Brazil K001106388 Yes
Calderon, C.E. [2833], Brazil K001106415 Yes
Calderon, C.E. [2846], Brazil K001106410 Yes
Calderon, C.E. [2829], Brazil K001106405 Yes
Trail, J.W.H. [1280], Brazil K001106398 Yes
Calderon, C.E. [2873], Brazil K001106365 Yes
Calderon, C.E. [2829], Brazil K001106406 Yes
Setgio, L. [303], Brazil K001106402 Yes
Calderon, C.E. [2833], Brazil K001106417 Yes
Calderon, C.E. [2846], Brazil K001106408 Yes
Calderon, C.E. [2282], Brazil K001106362 Yes
Calderon, C.E. [2282], Brazil K001106363 Yes
Stannard, B.L. [469], Venezuela K000743050 No
Guedes, J. [2754], Brazil K001106355 Yes
Calderon, E.C. [2818], Brazil K001106359 Yes
Calderon, C.E. [2910], Brazil K001106387 Yes
Kubitzki, K. [79-236], Brazil K001106354 Yes
Spruce, R. [420], Brazil K001106400 Yes
Martinelli, G. [9252], Brazil K001106352 Yes
Calderon, C.E. [2829], Brazil K001106404 Yes
Calderon, C.E. [2846], Brazil K001106407 Yes
Calderon, C.E. [2754], Brazil K001106356 Yes
Hatschbach, G. [20546], Brazil K001106396 Yes
Calderon, C.E. [2818], Brazil K001106357 Yes
Pereira, B.A.S. [2091], Brazil K001106386 Yes
Calderon, C.E. [2910], Brazil K001106389 Yes
Kubitzki, K. [79/236], Brazil K001106353 Yes
Burchell [8970], Brazil K001106397 Yes
Calderon, C.E. [2355], Brazil K001106351 Yes
Calderon, C.E. [2833], Brazil K001106418 Yes
Irwin, H.S. [6714], Brazil K001106390 Yes
Calderon, C.E. [2818], Brazil K001106360 Yes
Irwin, H.S. [6714], Brazil K001106391 Yes
Calderon, C.E. [2818], Brazil K001106361 Yes
Calderón, E. [2355], Brazil K001106350 Yes
Calderon, C.E. [2818], Brazil K001106358 Yes
Eiten, G. [10595], Brazil K001106394 Yes
Calderon, C.E. [2813], Brazil K001106412 Yes
Ule, E. [9147], Brazil K001106392 Yes
Spruce, R. [420], Brazil K001106401 Yes
Martinelli, G. [12008], Brazil K001106347 Yes
Acevedo-Rodriguez, P. [8334], Brazil K001106403 Yes
Eiten, G. [10595], Brazil K001106393 Yes
Hatschbach, G. [20546], Brazil K001106395 Yes

First published in J. Phys. Chim. Hist. Nat. Arts 95: 150 (1822)

Accepted by

  • Govaerts, R.H.A. (2011). World checklist of selected plant families published update Facilitated by the Trustees of the Royal Botanic Gardens, Kew.

  • GrassBase - The Online World Grass Flora

    http://creativecommons.org/licenses/by-nc-sa/3.0

  • Herbarium Catalogue Specimens

    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

  • Kew Backbone Distributions

    The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

  • Kew Names and Taxonomic Backbone

    The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

  • Kew Science Photographs

    Copyright applied to individual images