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This species is accepted, and its native range is S. Somalia to S. Africa, Madagascar.
A specimen from Kew's Herbarium

[FSOM]

M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS

Morphology General Habit
Shrub or small tree, up to c. 5 m tall; young branches with o without lenticels
Morphology Leaves
Leaf-blades narrowly elliptic to suborbicular, 2–10 x 1–4 cm, rounded to acute at the apex, cuneate to rounded at the base, glabrous to pubescent
Morphology Reproductive morphology Flowers
Flowers 4-merous, in axillary inflorescences
Morphology Reproductive morphology Flowers Calyx
Calyx-lobes broadly ovate to suborbicular, 1.7–2.5 mm long
Morphology Reproductive morphology Flowers Corolla
Corolla 5–8 mm long, hairy in throat
Morphology Reproductive morphology Flowers Gynoecium Pistil
Pistil 4–6 mm long, with ovary hairy above
Morphology Reproductive morphology Fruits
Fruit orange to yellow, 2–8 cm in diam., with a thick wall, several-seeded.
Distribution
S1 Kenya and southwards to South Africa and Madagascar
Ecology
Altitude c. 200 m.

[FZ]

Loganiaceae, A. J. M. Leeuwenberg. Flora Zambesiaca 7:1. 1983

Morphology General Habit
Often many–stemmed and much–branched shrub or small tree, 1·50–10(20) m. high, deciduous.
Morphology Trunk
Trunk 7–60 cm. in diam., mostly branched from low down.
Morphology General Bark
Bark mostly pale grey or greyish–white, smooth; wood whitish.
Morphology Branches
Branches pale grey to dark brown, powdery or not, lenticellate or not, smooth; branchlets glabrous or pubescent, terete.
Morphology Leaves
Leaves: petiole glabrous or pubescent, often short, 1–5 mm. long; lamina shining and dark green above, paler beneath, coriaceous, very variable in shape and size, suborbicular, elliptic, narrowly elliptic, obovate or narrowly obovate, (1)1·5–4(5) times as long as wide, 2–10(15·5) x 1·4(5·5) cm., rounded, or especially in Madagascar and near the coast in Mozambique and Natal and furthermore elsewhere in shade branches acute or acuminate at the apex, cuneate or rounded at the base, glabrous or pubescent on both sides; one or two pairs of distinct secondary veins from or from above the base curved along the margin and often a faint submarginal pair; tertiary venation reticulate, not or slightly prominent above.
Morphology Reproductive morphology Inflorescences
Inflorescences axillary or ramiflorous, usually several together, very short and nearly fasciculate, 1 x 1·2 x 1·5 cm., 1·2 x branched, few–flowered. Peduncle, branches, and pedicels short or very short.
Morphology Reproductive morphology Flowers
Flowers 4–merous.
Morphology Reproductive morphology Flowers Calyx
Sepals pale green, free or nearly so, subequal, the inner slightly smaller, ovate, broadly ovate or suborbicular, 1–1·6 times as long as wide, 1·7–2·5(3) x 1·5–2·2 mm., rounded or obtuse at the apex, ciliate, glabrous or with some minute appressed hairs at the base inside, without colleters.
Morphology Reproductive morphology Flowers Corolla
Corolla in the mature bud (1·8)2·8–3·3(3·6) times as long as the calyx, (4·5)5–8 mm. long, white or greenish–yellow, glabrous outside, inside with a brush–like ring of white lanate hairs in the throat and just on the base of the lobes; tube cylindrical or nearly so (1)1·2–1·8(2·2) times as long as the calyx, 0·7–1·7 x as long as the lobes, 2·5–4·5(5) mm. long, 1·5–2·8 mm. wide at the throat; lobes thick, narrowly triangular, 1·5–3 times as long as wide, 2–3·6 x 1–1·9 mm., acute or subacute, spreading.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens hardly exserted; filaments extremely short, inserted at the mouth of the corolla tube; anthers oblong, about twice as long as wide, 1·2–1·8 x 0·6–0·8 mm., glabrous.
Morphology Reproductive morphology Flowers Gynoecium Pistil
Pistil hirto–pilose in the middle, (3·5)4–6 mm. long; ovary narrowly ovoid or oblong, 1·5–3 x 0·8–1·5 mm., hirto–pilose at the very apex, further glabrous, often with a disc–like base, gradually narrowed into the style, 2–celled; style thick (2)2·5–4 mm. long, at the base hairy like the ovary at the apex; stigma capitate.
Morphology Reproductive morphology Flowers Gynoecium Ovary Ovules
In each cell 5–30 ovules.
Morphology Reproductive morphology Fruits
Fruit orange or yellow, nearly mature bluish–green, large or sometimes rather small, hard when not small, globose, 2–8(10) cm. in diam., with about 5–50 seeds, with somewhat granular skin, slightly shining. Wall mostly thick, (1)2–4 mm. thick, thicker above the pedicel, brittle in mature fruits, hard and not broken by hand when nearly mature and/or dry. Pulp orange, slimy, edible.
Morphology Reproductive morphology Seeds
Seeds pale ochraceous, flattened or not, often more or less plano–convex, obliquely ovate, elliptic, or tetrahedral, usually irregularly curved, 1–4–1–8 times as long as wide, 11–25 x 6–18 x 5–8 mm., with thick very short erect hairs, rather rough.

[FTEA]

Loganiaceae, E. A. Bruce & J. Lewis. Flora of Tropical East Africa. 1960

Morphology General Habit
Shrub or small much-branched tree, 2–12 m. high, with spreading branches and greyish to dark brown branchlets bearing round pale lenticels; young branchlets glabrous to pubescent, often densely covered with conspicuous elongate lenticels.
Morphology Leaves
Leaves subsessile, ± membranous to coriaceous, obovate to elliptic, very rarely more than twice as long as broad, 1.5–7 cm. long, 1–5 cm. wide, rounded or subacute, cuneate or rounded at the base, pubescent to glabrous and shining.
Morphology Reproductive morphology Inflorescences
Cyme axillary, shortly pedunculate or subsessile, pubescent or glabrous.
Morphology Reproductive morphology Flowers
Flowers 4-merous.
Morphology Reproductive morphology Flowers Calyx
Calyx-lobes suborbicular, about 2 mm. long, ciliate.
Morphology Reproductive morphology Flowers Corolla
Corolla whitish or yellowish; tube cylindric, about 3.5 mm. long, hirsute at the throat; lobes very thick and cucullate, oblong, subacute.
Morphology Reproductive morphology Flowers Androecium Stamens Anthers
Anthers sessile in the corolla-throat.
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary elongate-ovoid; style hirsute at the base.
Morphology Reproductive morphology Fruits
Fruit 2.5–7 cm. diameter, globose; rind hard, yellow, rarely black.
Morphology Reproductive morphology Seeds
Seeds about 7–12, tetrahedral, fairly thick, immersed in edible pulp.

Native to:

Angola, Botswana, Caprivi Strip, Kenya, KwaZulu-Natal, Madagascar, Malawi, Mozambique, Northern Provinces, Somalia, Swaziland, Tanzania, Zambia, Zimbabwe

Strychnos madagascariensis Poir. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Jan 1, 1999 Antilahimena, P. [218], Madagascar K000211271
Jan 1, 1996 Edmondson, N.Z. [63], Madagascar K000211272
Jan 28, 1995 Leeuwenberg, A.J.M. [14431], Madagascar K000211263
Jan 1, 1992 Labat, J.-.N. [2222], Madagascar K000211260
Jan 1, 1992 Labat, J.-.N. [2203], Madagascar K000211268
Jan 1, 1991 Leeuwenberg, A.J.M. [14167], Madagascar K000211259
Jan 1, 1991 Leeuwenberg, A.J.M. [14171], Madagascar K000211261
Jan 1, 1990 Dumetz, N. [1101], Madagascar K000211277
Jan 1, 1990 McPherson, G. [14692], Madagascar K000211275
Jan 1, 1989 Phillipson, P.B. [1708], Madagascar K000211285
Jan 1, 1989 Nicoll, M. [163], Madagascar K000211284
Jan 1, 1989 Nicoll, M. [350], Madagascar K000211283
Jan 1, 1974 Bernardi, L. [11902], Madagascar K000211278
Jan 1, 1974 Richard, A. [012], Madagascar K000211289
Jan 1, 1974 Bernardi, L. [11198], Madagascar K000211269
Jan 1, 1971 Yalala, A. M. [453], Botswana K000489701
Jan 1, 1967 Holst, C. [2420], Tanzania K000426637 isotype
Jan 1, 1967 Wakefield, T. [s.n.], Kenya K000426638 holotype
Jan 1, 1967 ken [726], South Africa K000426767
Jan 1, 1967 Wood, J.M. [5624], South Africa K000426768 lectotype
Jan 1, 1967 Gerrard [1421], South Africa K000426769 paratype
Jan 1, 1967 Kirk, J. [s.n.], Mozambique K000426792 paratype
Jan 1, 1967 Kirk, J. [368], Mozambique K000426800 lectotype
Jan 1, 1967 Burton [s.n.], Mozambique K000426801 paratype
Jan 1, 1967 Waller, H. [s.n.], Malawi K000426802 paratype
Jan 1, 1967 Forbes. [s.n.], Mozambique K000426803 holotype
Jan 1, 1967 Baron, R. [2419], Madagascar K000211267
Jan 1, 1967 Wood, J.M. [1777], South Africa K000426766 paratype
Jan 1, 1967 Baron, R. [4648], Madagascar K000426775 holotype
Jan 1, 1967 Kirk, J. [s.n.], Mozambique K000426793 paratype
Jan 1, 1967 Gerrard [1421], South Africa K000426770 paratype
Jan 1, 1967 Pervillé, A. [433], Madagascar K000211266
Jan 1, 1967 Capuron, R. [24336], Madagascar K000211281
Jan 1, 1967 Meller [s.n.], Madagascar K000211256
Jan 1, 1967 Perrier de la Bâthie, H. [s.n.], Madagascar K000211257
Jan 1, 1967 Pervillé, A. [696], Madagascar K000211265
Jan 1, 1967 Capuron, R. [24336], Madagascar K000211280
Jan 1, 1967 Bernier [s.n.], Madagascar K000211282
Jan 1, 1967 Greenway, P.J. [4851], Tanzania K000426636 isotype
Jan 1, 1967 Baron, R. [4648], Madagascar K000426776 isotype
Jan 1, 1967 Son, F. van [28860], Botswana K000489700
Jan 1, 1967 Baron, R. [2419], Madagascar K000211258
Jan 1, 1967 Rodin, R.J. [4510], Zambia K000197251
Jan 1, 1967 Humbert, H. [24376], Madagascar K000211279
Milne-Redhead, E. [7283], Tanzania 11156.000
White, F. [1990], Zambia 17673.000
Drummond, R.B. [4077], Kenya 6563.000
Richards, H.M. [24787], Tanzania 7737.000
Milne-Redhead, E. [8552], Tanzania 9873.000
Leeuwenberg, A.J.M. [14403], Madagascar K000211262
Miller, J.S. [4525], Madagascar K000211276
Smith, P.A. [4237], Botswana K000489698
Dorr, L.J. [3021], Madagascar K000211286
Zjhra, M.L. [229], Madagascar K000211273
Gautier, L. [3781], Madagascar K000211270
Smith, P.A. [2055], Botswana K000489699
Lorence, D. [1913], Madagascar K000211287
Phillipson, P.B. [2313], Madagascar K000211274
Rabevohitra, R. [1813], Madagascar K000211264
Richard, A. [139], Madagascar K000211288

First published in J.B.A.M.de Lamarck, Encycl. 8: 696 (1808)

Accepted by

  • Exell, A.W. & Wild, H. (eds.) (1960). Flora Zambesiaca 1(1): 1-336. Royal Botanic Gardens, Kew.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Mannheimer, C.A. & Curtis, B.A. (eds.) (2009). Le Roux and Müller's field guide to the trees and shrubs of Namibia, rev. ed.: 1-525. Macmillan Education Namibia, Windhoek.
  • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
  • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.

Literature

Kew Backbone Distributions

  • Bruce, E.A. & Lewis, J. (1960). Flora of Tropical East Africa, Loganiaceae: 1-47.
  • Exell, A.W. & Wild, H. (eds.) (1960). Flora Zambesiaca 1(1): 1-336. Royal Botanic Gardens, Kew.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Mannheimer, C.A. & Curtis, B.A. (eds.) (2009). Le Roux and Müller's field guide to the trees and shrubs of Namibia, rev. ed.: 1-525. Macmillan Education Namibia, Windhoek.
  • Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
  • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.

Flora of Somalia

  • Flora Somalia, Vol 3, (2006) Author: by M. Thulin [updated by M. Thulin 2008]

Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Somalia
Flora of Somalia
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0