- Family:
- Lamiaceae Martinov
Isodon (Schrad. ex Benth.) Spach

[LKGF]
Harley, R.M. et al. (2004). Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Springer-Verlag, Berlin, Heidelberg, New York.
- Morphology General Habit
- Shrubs or herbs
- Morphology Leaves
- Leaves simple, opposite, rarely ternate
- Morphology Reproductive morphology Inflorescences
- Inflorescence thyrsoid with cymes pedunculate, bracteate, bracteolate, 3- to several-flowered
- Morphology Reproductive morphology Inflorescences Bracts
- Bracts persistent, not coloured, smaller than leaves
- Morphology Reproductive morphology Inflorescences Bracteoles
- Bracteoles small, usually linear
- Morphology Reproductive morphology Flowers Calyx
- Calyx campanulate, funnel-shaped, or tubular and sometimes slightly constricted or rarely curved at throat, actinomorphic or 2-lipped, 5-lobed (3/2 or regular), lobes lanceolate to deltoid, posterior never decurrent, throat open, glabrous
- Morphology Reproductive morphology Flowers Corolla
- Corolla strongly 2-lipped, 5-lobed (4/1), white, blue, pink or purplish, posterior lip ascending or reflexed, 4-lobed with lobes equal or rarely median more shallowly lobed, anterior lip horizontal, cucullate, corolla-tube straight or geniculate, saccate
- Morphology Reproductive morphology Flowers Androecium Stamens
- Stamens 4, held within, or more rarely exceeding the anterior lip, posterior pair attached near base of corolla or between base and midpoint, usually with hairs, anterior pair attached between midpoint of corolla and throat, often with hairs
- Morphology Reproductive morphology Flowers Gynoecium Stigma
- Stigma-lobes diverging
- Morphology Reproductive morphology Flowers Disc
- Disc 4-lobed with anterior lobe slightly larger, lobes alternating with nutlets
- Morphology Reproductive morphology Fruits
- Nutlets ovoid, rarely conical with a flattened base, glabrous, sometimes with hairs or glands, mucilaginous or not, areole small, inconspicuous.
- Note
- 2n = 24, 42. Around 100 species
- Distribution
- Tropical Asia with two species in Africa.
[KBu]
Suddee, S., A. J. Paton, & Parnell, J. (2004). A Taxonomic Revision of Tribe Ocimeae Dumort. (Lamiaceae) in Continental South East Asia I. Introduction, Hyptidinae & Hanceolinae. Kew Bulletin, 59(3), 337-378. doi:10.2307/4110949
- Morphology General Habit
- Annual or perennial herbs or undershrubs
- Morphology Leaves
- Leaves membranous to chartaceous, usually serrate or sometimes serrate-crenate, lower pairs petiolate, upper pairs usually sessile
- Morphology Reproductive morphology Flowers Androecium Stamens
- Stamens exserted or included in anterior corolla lip; insertion of anterior pair varying from above the middle of corolla tube to the base of anterior corolla lip, glabrous or only slightly pubescent at base; posterior pair inserted near the base of corolla tube, sparsely or densely villous at base
- Morphology Reproductive morphology Flowers Calyx
- Calyx campanulate or tubular-campanulate, straight or declinate, subequally 5-toothed or bilabiate with posterior lip 3-lobed and anterior lip 2-lobed; tube 10-nerved, usually twice or more as long as teeth, without spur at anterior base
- Morphology Reproductive morphology Flowers Corolla
- Corolla with posterior lip subequally 4-lobed, usually pubescent with sessile glands on back; anterior lip concave or flattened, glabrous inside, usually pubescent with sessile glands outside; tube tubular, straight or declinate, usually gibbous on posterior side near base
- Morphology Reproductive morphology Flowers Disc
- Disc with anterior side well or slightly developed
- Morphology Reproductive morphology Flowers Gynoecium Ovary
- Ovary glabrous
- Morphology Reproductive morphology Flowers Gynoecium Style
- Style shortly bifid with subequal branches, shorter or longer than stamens
- Morphology Reproductive morphology Fruits
- Nutlets oblong, ovoid or ellipsoid, smooth or minutely tuberculate, producing mucilage when wet or not.
- Morphology Reproductive morphology Inflorescences
- Inflorescence terminal and axillary, usually branched, forming a small or large panicle; cymes sessile or pedunculate, lax or dense, few- to many-flowered; bracts with lower pairs similar to leaves, gradually reduced in size upwards, persistent; bracteoles present, short, caducous or persistent
- Morphology Stem
- Stems sometimes woody at base, quadrangular or round- quadrangular, glabrous or pubescent and usually hollow inside
- Note
- Plectranthus L'Her. sect. Pyramidium Benth. was first published in Wall., Pl. Asiat. Rar. 2: 17 (1830), not in Labiat. Gen. Spec.: 44 (1832). Rabdosiella calycinus (Benth.) Codd, the type species of Rabdosiella, is actually a Plectranthus (Ryding 1993b).
- Type
- Lectotype species: Isodon rugosus (Wall. ex Benth.) Codd (Plectranthus rugosus Wall. ex Benth.).
[FTEA]
Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Isodon, RM Polhill. Flora of Tropical East Africa. 2009
- Morphology General Habit
- Herbs or shrubs
- Morphology Stem
- Stems quadrangular, with simple sometimes septate hairs, glandular hairs and sessile glands
- Morphology Leaves
- Leaves undivided, opposite, rarely ternate, usually petiolate
- Morphology Reproductive morphology Inflorescences
- Inflorescences thyrsoid, extended with cymes pedunculate, dichasial lax, elsewhere sometimes more congested; bracts leaf-like, smaller above; bracteoles present
- Morphology Reproductive morphology Flowers Calyx
- Calyx campanulate to tubular, accrescent, sometimes becoming slightly constricted and curved at throat, subequally 5-toothed, sometimes slightly 2-lipped, the posterior lip with 3 teeth, the anterior with 2 teeth
- Morphology Reproductive morphology Flowers Corolla
- Corolla white, blue, pink or purplish, 2-lipped; tube straight or geniculate, ± saccate to spurred at base, ± parallelsided distally; posterior lip short, recurved or reflexed, 4-lobed; anterior lip flat to boat-shaped, ± horizontal
- Morphology Reproductive morphology Flowers Androecium Stamens
- Stamens 4, declinate, posterior pair attached near base of corolla, anterior pair attached in upper part of corolla tube
- Morphology Reproductive morphology Flowers Disc
- Disk 4-lobed, with anterior lobe slightly larger
- Morphology Reproductive morphology Flowers Gynoecium Ovary
- Ovary deeply 4-lobed; style gynobasic, exserted, bifid with lobes short, equal and diverging
- Morphology Reproductive morphology Fruits
- Nutlets ovoid to ellipsoid or rarely conical, glabrous, mucilaginous when wetted or not; attachment scar basal, small.
Native to:
Afghanistan, Amur, Angola, Assam, Bangladesh, Burundi, Cameroon, China North-Central, China South-Central, China Southeast, East Himalaya, Ethiopia, Gulf of Guinea Is., India, Japan, Jawa, Kenya, Khabarovsk, Korea, Laos, Lesser Sunda Is., Malawi, Malaya, Manchuria, Myanmar, Nepal, Nigeria, Oman, Pakistan, Philippines, Primorye, Rwanda, Saudi Arabia, Sierra Leone, Sri Lanka, Sudan, Sulawesi, Sumatera, Taiwan, Tanzania, Thailand, Tibet, Uganda, Vietnam, West Himalaya, Zambia, Zaïre, Zimbabwe
- Isodon adenanthus (Diels) Kudô
- Isodon adenolomus (Hand.-Mazz.) H.Hara
- Isodon × akitaensis Fujiw., Yu.Abe & Y.Matsuda
- Isodon albopilosus (C.Y.Wu & H.W.Li) H.Hara
- Isodon amethystoides (Benth.) H.Hara
- Isodon angustifolius (Dunn) Kudô
- Isodon × arakii Murata
- Isodon assamicus (Mukerjee) H.Hara
- Isodon atroruber R.A.Clement
- Isodon aurantiacus Y.P.Chen & C.L.Xiang
- Isodon barbeyanus (H.Lév.) H.W.Li
- Isodon brevicalcaratus (C.Y.Wu & H.W.Li) H.Hara
- Isodon brevifolius (Hand.-Mazz.) H.W.Li
- Isodon bulleyanus (Diels) Kudô
- Isodon calcicola (Hand.-Mazz.) H.Hara
- Isodon capillipes (Benth.) H.Hara
- Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
- Isodon colaniae H.Hara ex Suddee & A.J.Paton
- Isodon dawoensis (Hand.-Mazz.) H.Hara
- Isodon delavayi C.L.Xiang & Y.P.Chen
- Isodon dhankutanus Murata
- Isodon effusus (Maxim.) H.Hara
- Isodon enanderianus (Hand.-Mazz.) H.W.Li
- Isodon eriocalyx (Dunn) Kudô
- Isodon excisoides (Y.Z.Sun ex C.H.Hu) H.Hara
- Isodon excisus (Maxim.) Kudô
- Isodon flabelliformis (C.Y.Wu) H.Hara
- Isodon flavidus (Hand.-Mazz.) H.Hara
- Isodon flexicaulis (C.Y.Wu & H.W.Li) H.Hara
- Isodon forrestii (Diels) Kudô
- Isodon gesneroides (J.Sinclair) H.Hara
- Isodon gibbosus (C.Y.Wu & H.W.Li) H.Hara
- Isodon glutinosus (C.Y.Wu & H.W.Li) H.Hara
- Isodon grandifolius (Hand.-Mazz.) H.Hara
- Isodon grosseserratus (Dunn) Kudô
- Isodon henryi (Hemsl.) Kudô
- Isodon hirtellus (Hand.-Mazz.) H.Hara
- Isodon hispidus (Benth.) Murata
- Isodon × inamii Murata
- Isodon inflexus (Thunb.) Kudô
- Isodon interruptus (C.Y.Wu & H.W.Li) H.Hara
- Isodon irroratus (Forrest ex Diels) Kudô
- Isodon japonicus (Burm.f.) H.Hara
- Isodon kurzii (Prain) H.Hara
- Isodon latifolius (C.Y.Wu & H.W.Li) H.Hara
- Isodon leucophyllus (Dunn) Kudô
- Isodon liangshanicus (C.Y.Wu & H.W.Li) H.Hara
- Isodon lihsienensis (C.Y.Wu & H.W.Li) H.Hara
- Isodon longitubus (Miq.) Kudô
- Isodon lophanthoides (Buch.-Ham. ex D.Don) H.Hara
- Isodon loxothyrsus (Hand.-Mazz.) H.Hara
- Isodon lungshengensis (C.Y.Wu & H.W.Li) H.Hara
- Isodon macrocalyx (Dunn) Kudô
- Isodon macrophyllus (Migo) H.Hara
- Isodon medilungensis (C.Y.Wu & H.W.Li) H.Hara
- Isodon meeboldii (W.W.Sm.) Suddee
- Isodon megathyrsus (Diels) H.Hara
- Isodon melissoides (Benth.) H.Hara
- Isodon mucronatus (C.Y.Wu & H.W.Li) H.Hara
- Isodon muliensis (W.W.Sm.) Kudô
- Isodon myriocladus C.Chen
- Isodon namikawanus Murata
- Isodon nervosus (Hemsl.) Kudô
- Isodon nigrescens (Benth.) H.Hara
- Isodon nilgherricus (Benth.) H.Hara
- Isodon × ohwii Okuyama
- Isodon oreophilus (Diels) A.J.Paton & Ryding
- Isodon oresbius (W.W.Sm.) Kudô
- Isodon pantadenius (Hand.-Mazz.) H.W.Li
- Isodon parvifolius (Batalin) H.Hara
- Isodon pharicus (Prain) Murata
- Isodon phulchokiensis (Murata) H.Hara
- Isodon phyllopodus (Diels) Kudô
- Isodon phyllostachys (Diels) Kudô
- Isodon pleiophyllus (Diels) Kudô
- Isodon purpurescens Sunil, Naveen Kum. & Ratheesh
- Isodon racemosus (Hemsl.) Murata
- Isodon ramosissimus (Hook.f.) Codd
- Isodon repens (Wall. ex Benth.) Murata
- Isodon rivularis (Wight ex Hook.f.) H.Hara
- Isodon rosthornii (Diels) Kudô
- Isodon rubescens (Hemsl.) H.Hara
- Isodon rugosiformis (Hand.-Mazz.) H.Hara
- Isodon rugosus (Wall. ex Benth.) Codd
- Isodon schimperi (Vatke) J.K.Morton
- Isodon scoparius (C.Y.Wu & H.W.Li) H.Hara
- Isodon scrophularioides (Wall. ex Benth.) Murata
- Isodon sculponeatus (Vaniot) Kudô
- Isodon secundiflorus (C.Y.Wu) H.Hara
- Isodon serra (Maxim.) Kudô
- Isodon setschwanensis (Hand.-Mazz.) H.Hara
- Isodon shikokianus (Makino) H.Hara
- Isodon silvaticus (C.Y.Wu & H.W.Li) H.W.Li
- Isodon smithianus (Hand.-Mazz.) H.Hara
- Isodon × suzukii Okuyama
- Isodon tenuifolius (W.W.Sm.) Kudô
- Isodon ternifolius (D.Don) Kudô
- Isodon teysmannii (Miq.) H.W.Li
- Isodon × togashii Okuyama
- Isodon trichocarpus (Maxim.) Kudô
- Isodon umbrosus (Maxim.) H.Hara
- Isodon villosus Y.P.Chen & H.Peng
- Isodon walkeri (Arn.) H.Hara
- Isodon wardii (C.Marquand & Airy Shaw) H.Hara
- Isodon websteri (Hemsl.) Kudô
- Isodon weisiensis (C.Y.Wu) H.Hara
- Isodon wightii (Benth.) H.Hara
- Isodon wikstroemioides (Hand.-Mazz.) H.Hara
- Isodon wui C.L.Xiang & E.D.Liu
- Isodon xerophilus (C.Y.Wu & H.W.Li) H.Hara
- Isodon yuennanensis (Hand.-Mazz.) H.Hara
Isodon (Schrad. ex Benth.) Spach appears in other Kew resources:
First published in Hist. Nat. Vég. 9: 162 (1838)
Accepted by
- Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
Literature
Flora of Tropical East Africa
- Bothalia 15: 7–9 (1984)
- Hist. Nat. Vég. Phan. 9: 162 (1840)
- J. Arn. Arbor. 69: 289–400 (1988)
- K.B. 53: 723–731 (1998)
- K.B. 58: 487 (2003)
- Kubitzki, Fam. Gen. Vasc. Pl. 7: 256 (2004)
- Pl. Syst. Evol. 185: 91–97 (1993)
- Taxon 17: 239 (1968)
Lamiaceae Key Genus Fact Sheets
- Amethystanthus Nakai, Bot. Mag. (Tokyo) 48: 785 (1934).
- Dielsia Kudô, Mem. Fac. Sci. Taihoku Imp. Univ. 2: 143 (1929), nom. illeg.
- Harley RM, Atkins S, Budantsev AL, Cantino PD, Conn BJ, Grayer R, Harley MM, de Kok RPJ, Krestovskaja T, Morales R, Paton AJ, Ryding O, and Upson T. 2004. Labiatae, in The Families and Genera of Vascular Plants (K. Kubitzki, ed. in chief) VI: 167-275. Sp
- Homalocheilos J.K.Morton, J. Linn. Soc., Bot. 58: 249 (1962).
- Isodon (Benth.) Schrader ex Spach, Hist. Nat. Vég. 9: 162 (1840)
- Li, J. Arnold Arbor. 69: 289-400 (1988), rev.
- Paton & Ryding, Kew Bull. 53: 723-731 (1998).
- Plectranthus L'Hér. sect. Amethystoides Schrad. ex Benth. (1832).
- Plectranthus L'Hér. sect. Isodon Schrad. ex Benth. (1832).
- Plectranthus L'Hér. sect. Melissoides Schrad. ex Benth. (1832).
- Plectranthus L'Hér. sect. Pyramidatum Schrad. ex Benth. (1832).
- Rabdosiella Codd, Bothalia 15: 9 (1984).
- Skapanthus C.Y.Wu & H.W.Li, Acta Phytotax. Sin. 13: 77 (1975).
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Flora of Tropical East Africa
Flora of Tropical East Africa
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Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0
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Kew Bulletin
Kew Bulletin
http://creativecommons.org/licenses/by-nc-sa/3.0
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Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0
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Lamiaceae Key Genus Fact Sheets
Nina Davies, Gemma Bramley and Don Kirkup, Royal Botanic Gardens, Kew
http://creativecommons.org/licenses/by/3.0