- Family:
- Ceratophyllaceae Gray
- Genus:
- Ceratophyllum L.
Ceratophyllum submersum L.

[FTEA]
Ceratophyllaceae, C.M. Wilmot-Dear. Flora of Tropical East Africa. 1985
- Morphology General Habit
- Very similar to C. demersum in form, but usually more delicate and filamentous in appearance, differing as follows.
- Morphology Leaves
- Leaves usually lighter green, 2–4 cm. long, branched dichotomously 3–4 times, sometimes one part of leaf branched more than another, rarely a few leaves of whorl only twice branched, usually rather fine, 0.1–0.3 mm. in diameter, but lower 1–2 segments sometimes swollen and up to 2.5 mm. across, marginal spiny teeth always minute, less than 0.1 or 0.2 mm., on lower 2 segments few or absent, few or more rarely numerous on final 2 segments.
- sex Male
- Male flowers up to 2 mm. in diameter; mature anthers 0.6–0.8 mm. long, 0.4–0.7 mm. wide (length 1–1.5 times width).
- Morphology Reproductive morphology Flowers
- Female perianth-segments usually longer, (1.5–)1.8–2 mm. long, 0.1–0.3 mm. wide, and relatively narrower (length (5–)7–10 times width). Male flowers up to 2 mm. in diameter; mature anthers 0.6–0.8 mm. long, 0.4–0.7 mm. wide (length 1–1.5 times width).
- sex Female
- Female perianth-segments usually longer, (1.5–)1.8–2 mm. long, 0.1–0.3 mm. wide, and relatively narrower (length (5–)7–10 times width).
- Morphology Reproductive morphology Fruits
- Mature fruit with apical spine less than 1 up to 9 mm.; surface sometimes with few large warts or spines.
[FZ]
Ceratophyllaceae, C. M. Wilmot-Dear. Flora Zambesiaca 9:6. 1991
- Morphology General Habit
- Aquatic herb similar to Ceratophyllum demersum, but usually more delicate, differing as follows: Leaves usually lighter green, 2–4 cm. long, branching dichotomously 3–4 times (rarely twice in some whorls), usually rather fine, 0.1–0.3 mm. thick, to 2.5 mm. thick in lower segments, marginal teeth few, inconspicuous, often absent from lower segments, up to 0.1–0.2 mm. long.
- sex Male
- Male flowers to 2 mm. in diam.; anthers 0.6–0.8 x 0.4–0.7 mm., often smaller and relatively broader than in C. demersum.
- Morphology Reproductive morphology Flowers
- Female perianth lobes (1.5) 1.8–2 x 0.1–0.3 mm., relatively narrower and usually longer than in C. demersum. Male flowers to 2 mm. in diam.; anthers 0.6–0.8 x 0.4–0.7 mm., often smaller and relatively broader than in C. demersum.
- sex Female
- Female perianth lobes (1.5) 1.8–2 x 0.1–0.3 mm., relatively narrower and usually longer than in C. demersum.
- Morphology Reproductive morphology Fruits
- Fruit usually markedly laterally flattened with a marginal rim or wing.
Native to:
Albania, Algeria, Altay, Austria, Baltic States, Bangladesh, Belarus, Belgium, Bulgaria, Cameroon, Central European Rus, Chad, Corse, Czechoslovakia, Denmark, East European Russia, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Kazakhstan, Kenya, Kirgizstan, Krym, Libya, Morocco, Netherlands, Nigeria, North Caucasus, Northwest European R, Oman, Palestine, Poland, Romania, Sicilia, South European Russi, Spain, Sweden, Switzerland, Tanzania, Tunisia, Turkey, Ukraine, Uzbekistan, West Siberia, Yugoslavia, Zaïre
Introduced into:
Finland
Ceratophyllum submersum L. appears in other Kew resources:
First published in Sp. Pl. ed. 2: 1409 (1763)
Accepted by
- Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
- Danin, A. & Fragman- Sapir, O. (2019). Flora of Israel Online http://flora.org.il/en/plants/.
- Dobignard, A. & Chatelain, C. (2011). Index synonymique de la flore d'Afrique du nord 3: 1-449. Éditions des conservatoire et jardin botaniques, Genève.
- Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
- Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
- Lazkov, G.A. & Sultanova, B.A. (2011). Checklist of vascular plants of Kyrgyzstan Norrlinia 24: 1-166.
- Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
- Mosti, S., Raffaelli, M. & Tardelli, M. (2012). Contributions to the flora of central-southern Dhofar (Sultanate of Oman) Webbia; Raccolta de Scritti Botanici 67: 65-91.
- Sheremetova, S.A., Ebel, A.L. & Buko, T.E. (2011). Supplement to the flora of Kemerovo region since 2001 till 2010 Turczaninowia 14(1): 65-74.
- Stace, C. (2019). New Flora of the British Isles ed. 4: 1-1266. C & M Floristics.
- Szalontai, B. & al. (2018). Molecular phylogenetic analysis of Ceratophyllum L. taxa: a new perspective Botanical Journal of the Linnean Society 188: 161-172.
- Takhtajan, A.L. (ed.) (2012). Takhtajan, A.L. (ed.) Konspekt Flora Kavkaza 3(2): 1-623. Editio Universitatis Petropolitanae.
Literature
Kew Backbone Distributions
- Danin, A. & Fragman- Sapir, O. (2019). Flora of Israel Online http://flora.org.il/en/plants/.
- Dobignard, A. & Chatelain, C. (2011). Index synonymique de la flore d'Afrique du nord 3: 1-449. Éditions des conservatoire et jardin botaniques, Genève.
- Jonsell, B. (ed.) (2001). Flora Nordica 2: 1-430. The Bergius Foundaton.
- Lazkov, G.A. & Sultanova, B.A. (2011). Checklist of vascular plants of Kyrgyzstan Norrlinia 24: 1-166.
- Mostaph, M.K. & Uddin, S.B. (2013). Dictionary of plant names of Bangladesh, Vasc. Pl.: 1-434. Janokalyan Prokashani, Chittagong, Bangladesh.
- Mosti, S., Raffaelli, M. & Tardelli, M. (2012). Contributions to the flora of central-southern Dhofar (Sultanate of Oman) Webbia; Raccolta de Scritti Botanici 67: 65-91.
- Sheremetova, S.A., Ebel, A.L. & Buko, T.E. (2011). Supplement to the flora of Kemerovo region since 2001 till 2010 Turczaninowia 14(1): 65-74.
- Stace, C. (2019). New Flora of the British Isles ed. 4: 1-1266. C & M Floristics.
- Takhtajan, A.L. (ed.) (2012). Takhtajan, A.L. (ed.) Konspekt Flora Kavkaza 3(2): 1-623. Editio Universitatis Petropolitanae.
Flora of Tropical East Africa
- A.D.Q. Agnew, Upland Kenya Wild Flowers p. 81 (1974).
- Hauman in Flore du Congo Belge et du Ruanda-Urundi, 2: 165 (1951).
- L., Sp. Pl., ed. 2: 1409 (1763).
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Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0
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Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0
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Herbarium Catalogue Specimens
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Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0
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Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0
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Kew Science Photographs
Copyright applied to individual images