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This species is accepted, and its native range is Tanzania to S. Africa.
Combretum imberbe Wawra

[FTEA]

Combretaceae, G. E. Wickens. Flora of Tropical East Africa. 1973

Morphology General Habit
Tree to 33 m. tall or a shrub, branches often becoming spiny; new growth densely rufous lepidote.
Morphology Leaves
Leaves opposite or subopposite; lamina silvery, papyraceous to subcoriaceous, narrowly elliptic to elliptic-oblong or elliptic-obovate, (2.5–)5–6(–8.5) cm. long, (l–)2–2.5(–3) cm. wide, apex obtuse or rounded and often mucronate, base obtuse to narrowly cuneate, densely silvery lepidote on both surfaces, otherwise glabrous, scales often contiguous or overlapping; lateral nerves 4–7 pairs, scarcely visible; petiole 4–10 mm. long, densely rufous lepidote.
Morphology Reproductive morphology Inflorescences
Inflorescence up to ± 10 cm. long, spikes often forming a terminal panicle by suppression of the upper leaves, or unbranched lateral spikes up to 5 cm. long.
Morphology Reproductive morphology Flowers
Flowers yellowish.
Morphology Reproductive morphology Flowers Receptacle
Lower and upper receptacles densely silvery or rufous lepidote, otherwise glabrous or nearly so.
Morphology Reproductive morphology Flowers Calyx
Sepals ovate-triangular.
Morphology Reproductive morphology Flowers Corolla
Petals obovate to spathulate, 1–1.2 mm. long, 0.4–0.6 mm. wide, glabrous.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamen-filaments 1.5–2 mm. long; anthers 0.3–0.4 mm. long.
Morphology Reproductive morphology Flowers Disc
Disk 2–2.5 mm. in diameter, margin densely tomentose.
Morphology Reproductive morphology Flowers Gynoecium Style
Style 2 mm. long, lower part densely glandular.
Morphology Reproductive morphology Fruits
Fruit (fig. 4/4, p. 15) subcircular to broadly ovate in outline, 1.3–1.8 cm. long, 1.3–1.8 cm. wide, fairly densely to densely silvery lepidote, otherwise glabrous or nearly so, apex pointed; apical peg up to 1 mm. long or absent; wings up to 7 mm. broad; stipe 2–3 mm. long.
Morphology Reproductive morphology Seeds Germination
Cotyledons 2, 15–18 mm. long, 15 mm. wide, with petioles 9–11 mm. long, arising at or below soil-level.
Morphology General Scales
Scales (fig. 1/4, p. 12) 120–300 μ in diameter, roughly circular, cells very numerous and small (marginal cells ± 40–100), divided by many radial and tangential walls; cells beneath the scales with conspicuous round papillae.
Habitat
Locally common in wooded grassland on alluvial soils and “black cotton” soils; 0–1000 m.
Distribution
T6 T8

[FZ]

Combretaceae, A. W. Exell. Flora Zambesiaca 4. 1978

Morphology General Habit
Tree up to 30 m. tall (usually 6–10 m.) or shrub; bark light-grey or whitish; branchlets often becoming spiny, glabrous.
Morphology Leaves
Leaves silvery, opposite or sub-opposite; lamina 2·5–8·5 x 1–3 cm., papyraceous to subcoriaceous, narrowly elliptic to elliptic-oblong, densely silvery-lepidote on both surfaces otherwise glabrous, scales often contiguous or overlapping, apex obtuse or rounded and often mucronate, base obtuse to narrowly cuneate; lateral nerves 4–7 pairs, slightly prominent on both surfaces, usually without domatia; petiole 4–10 mm. long, lepidote but otherwise glabrous or almost so.
Morphology Reproductive morphology Inflorescences
Inflorescences of spikes up to c. 10 cm. long, often forming a terminal panicle by suppression of the upper leaves, or unbranched lateral spikes up to 5 cm. long; rhachis lepidote, otherwise glabrous or sparsely puberulous; bracts absent or very caducous.
Morphology Reproductive morphology Flowers
Flowers sessile, yellowish, densely lepidote but otherwise glabrous or nearly so.
Morphology Reproductive morphology Flowers Receptacle
Lower receptacle 2·5(3) x 0·25–0·3 mm., densely silvery- or rufous-lepidote, otherwise glabrous or nearly so; upper receptacle little developed, almost flat when the flowers are mature, densely silvery- or rufous-lepidote, otherwise glabrous or nearly so.
Morphology Reproductive morphology Flowers Calyx
Sepals 1·5 x 1·5 mm., ovate-triangular.
Morphology Reproductive morphology Flowers Corolla
Petals 1–1·2 x 0·4–0·6 mm., obovate to spathulate, glabrous.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamen-filaments 1·5–2 mm. long; anthers 0·3–0·4 mm. long.
Morphology Reproductive morphology Flowers Disc
Disk 2–2·5 mm. in diam., margin densely tomentose.
Morphology Reproductive morphology Fruits
Fruit 1·3–1·8 cm., subcircular to broadly ovate in outline, fairly densely to densely silvery-lepidote, apex pointed, apical peg very short (up to 0·5 mm.) to medium (1 mm.) or absent, wings up to 7 mm. broad, stipe 2–3 mm. long.
Morphology Reproductive morphology Seeds Cotyledons
Cotyledons 2, 15–18 x 15 mm., subcircular, with petioles 9–11 mm. long arising at or below soil level.
Morphology General Scales
Scales 120–300µ in diam., roughly circular, cells very numerous and small (marginal cells c. 40–100), divided as in the sectional description; cells beneath the scales with conspicuous round papillae.

Native to:

Angola, Botswana, Caprivi Strip, KwaZulu-Natal, Malawi, Mozambique, Namibia, Northern Provinces, Swaziland, Tanzania, Zambia, Zimbabwe

English
Bastard yellow wood, Elephant trunk, Ivory tree, Large fruited bushwillow, Large fruited combretum, Leadwood

Combretum imberbe Wawra appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Sep 23, 2008 Crawford, F. [59], Namibia K000450440
Jan 1, 1967 Kirk, J. [s.n.], Mozambique K000226544 holotype
Jan 1, 1961 Welwitsch [4372], Angola K000226542 Unknown type material
Jan 1, 1961 Welwitsch [4283], Angola K000226543 Unknown type material
Jan 1, 1960 Marloth [1264], Namibia K000226564 Unknown type material
Smith, P.A. [294], Botswana K000489040
Williams, D.T. [64], Botswana K000489041
Long, D.G. [318], Botswana K000489047
Smuts, J.C. [1988], South Africa K000812514
Hoogh, R.J. de [247], Botswana K000489043
Brown, R.C. [7572], Botswana K000489046
Allen, A. [231], Botswana K000489045
Kelaole, E.T. [586], Botswana K000489042
Allen, A. [360], Botswana K000489044
Long, D.G. [238], Botswana K000489048

First published in Sitzungsber. Kaiserl. Akad. Wiss., Math.-Naturwiss. Cl. 38: 556 (1860)

Accepted by

  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
  • Jongkind, C.C.H. (2014). Notes on African Combretum Loefl. Species (Combretaceae) Adansonia, sér. 3, 36: 315-327.
  • Launert, E. (ed.) (1978). Flora Zambesiaca 4: 1-658. Royal Botanic Gardens, Kew.
  • Mannheimer, C.A. & Curtis, B.A. (eds.) (2009). Le Roux and Müller's field guide to the trees and shrubs of Namibia, rev. ed.: 1-525. Macmillan Education Namibia, Windhoek.
  • Wickens, G.E. (1973). Flora of Tropical East Africa, Combretaceae: 1-99.

Literature

Kew Backbone Distributions

  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.

Flora of Tropical East Africa

  • Engl. & Diels in Monographieen Afrikanischer Pflanzen-Familien und Gattungen 3: 14 (1899).
  • Exell in Kirkia 7: 171 (1970).
  • F. White, Forest Flora of Northern Rhodesia p. 284 (1962).
  • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 138 (1949).
  • Wawra in Sitz. Akad. Wiss. Wien, Math.-Nat. Klasse 38: 556 (1860).

Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Plants and People Africa
Common Names from Plants and People Africa http://www.plantsandpeopleafrica.com/
© Plants and People Africa http://www.plantsandpeopleafrica.com http://creativecommons.org/licenses/by/3.0/