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This species is accepted, and its native range is Tropical & S. Africa, Arabian Peninsula.
A specimen from Kew's Herbarium

[FTEA]

Combretaceae, G. E. Wickens. Flora of Tropical East Africa. 1973

Morphology General Habit
Generally a small tree 5–7 m. high, sometimes 10–17 m. high, sometimes shrubby; bark dark grey to black, rough, reticulately fissured (resembling crocodile skin); branchlets with bark peeling in grey fibrous strips.
Morphology Leaves
Leaves opposite (sometimes 3-verticillate in coppice regrowth); lamina generally coriaceous, sometimes papyraceous (variant “A”), narrowly elliptic or narrowly ovate-elliptic or ovate-triangular or broadly ovate-elliptic or obovate or obovate-elliptic, up to 21 cm. long and 12.5 cm. wide, apex acute or obtuse, base cuneate or rounded or cordate, typically pubescent above and densely grey tomentose beneath, sometimes indumentum a dark velvety brown on the old leaves and pinkish on the young leaves, some forms almost glabrous above and pubescent beneath or glabrous on both surfaces (except for the scales); scales silvery or reddish, contiguous or almost so, but often hidden by the indumentum; lateral nerves 6–12 pairs, the parallel reticulation usually very prominent beneath (but less so in apparently transitional forms); petiole usually 2–3 mm. long.
Morphology Reproductive morphology Inflorescences
Inflorescences of axillary spikes up to 7(–11) cm. long, occasionally forming panicles by suppression of the upper leaves; peduncles 1–2 cm. long, pubescent.
Morphology Reproductive morphology Flowers
Flowers yellow or greenish yellow, fragrant.
Morphology Reproductive morphology Flowers Receptacle
Lower receptacle tomentose; upper receptacle campanulate, 1.5–3 mm. long, 2–3 mm. in diameter, tomentose, lepidote.
Morphology Reproductive morphology Flowers Calyx
Sepals broadly deltate.
Morphology Reproductive morphology Flowers Corolla
Petals sometimes minute or absent, irregularly obovate-deltate to reniform, 0.5–1 mm. long and wide, apex ciliate.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamen-filaments 5–6 mm. long, inserted at the margin of the disk; anthers 1 mm. long.
Morphology Reproductive morphology Flowers Disc
Disk with a very short pilose free margin less than 0.5 mm. wide.
Morphology Reproductive morphology Flowers Gynoecium Style
Style 3–5 mm. long.
Morphology Reproductive morphology Fruits
Fruit (fig. 5/14, p. 16) subcircular to elliptic in outline, 1.3–2.3(–2.5) cm. long, 1.5–2.3(–2.5) cm. wide, pale straw coloured to yellow-brown, lepidote and tomentose (especially on the body) to nearly glabrous (except for the scales); wings papyraceous (especially variant “A”); apical peg up to 1 mm. long; stipe 2.5 mm. long.
Morphology Reproductive morphology Seeds Germination
Cotyledons 2, arising at or below soil-level.
Morphology General Scales
Scales (fig. 2/14, p. 13) discoid with marginal cells usually concave, ± (75–)90–120 (–130) μ in diameter, usually with 8 radial walls and typically 8 (rarely 0) tangential walls and sometimes with many to several extra radial walls; marginal cells 8–11 (–16); cell walls clear and usually very thick; cells usually opaque.
Habitat
Widespread throughout the wooded grassland and bushland areas of East Africa, often forming pure stands on hillsides; 30–2300 m.
Distribution
throughout the wooded grassland areas of tropical and southern Africaalso in the Yemen (the only African species found outside Africa) K1 K2 K3 K4 K5 K6 K7 T1 T2 T3 T4 T5 T6 T7

[FZ]

Combretaceae, A. W. Exell. Flora Zambesiaca 4. 1978

Morphology Reproductive morphology Flowers Receptacle
Lower receptacle 1·5–2 mm. long, tomentose; upper receptacle 1·3–3 x 2–3 mm., campanulate, tomentose.
Morphology Reproductive morphology Flowers Calyx
Sepals 0·5–1 mm. long, broadly deltate.
Morphology Reproductive morphology Flowers Corolla
Petals 0·5–1 x 0·5–1 mm., sometimes minute or absent, irregularly obovate-deltate to reniform, ciliate at the apex.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamen-filaments 5–6 mm. long, inserted at the edge of the disk; anthers 1 mm. long.
Morphology Reproductive morphology Flowers Disc
Disk with a very short pilose free margin less than 0·5 mm. long.
Morphology Reproductive morphology Flowers Gynoecium Style
Style 5 mm. long, not expanded at the apex.
Morphology Reproductive morphology Fruits
Fruit 1·5–2(2·5) x 1·5–2(2·5) cm., subcircular to elliptic in outline, lepidote and tomentose (especially on the body) to nearly glabrous (except for scales), apical peg up to 1 mm. long (often much shorter), wings 5–7 mm. broad, stipe 2–3(5) mm. long.
Morphology Reproductive morphology Seeds Cotyledons
Cotyledons 2, 2·5 x 2·5–3·5 cm., subcircular to transversely elliptic, arising at or below soil level; petioles 3·5–4 cm. long.
Morphology General Scales
Scales ± contiguous on the areolae of the lower surface of the leaf, c. 75(90)–120(130)µ in diam., ± circular with marginal cells usually concave, usually with 8 radial walls and typically 8 (rarely 0) tangential walls and sometimes with many to several extra radial walls; marginal cells 8–11(16); cell-walls clear and usually very thick; cells usually opaque.
Morphology General Habit
Semi-deciduous tree usually up to 8(17) m. high or shrub; crown rounded to flat-rounded; bark black-brown to grey-brown, rough, fissured; branchlets tomentose to almost glabrous with bark coming off in fibrous strips.
Morphology Leaves
Leaves opposite; lamina up to 14(16) x 9 cm., narrowly elliptic or narrowly ovate-elliptic or obovate or obovate-elliptic, typically densely grey-tomentose (often drying dark-velvety-brown) but almost glabrous in some forms, lepidote but scales often hidden by the indumentum, reticulation usually very prominent beneath (but less so in some apparently transitional forms), apex acute or obtuse and often mucronate or apiculate, base usually rounded to subcordate; lateral nerves 6–12 pairs; petiole usually 2–3 mm. long (up to 9 mm.) leaving after leaf-fall rather prominent circular projections on the stem.
Morphology Reproductive morphology Inflorescences
Inflorescences of axillary spikes up to 7(11) cm. long occasionally forming panicles by suppression of the upper leaves; rhachis tomentose; bracts 1–2 mm. long, filiform, caducous; peduncles 1–2 cm. long.
Morphology Reproductive morphology Flowers
Flowers sessile, yellow or greenish-yellow.

[FSOM]

M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS

Morphology General Habit
Tree or shrub; vegetative parts glabrous to tomentose
Morphology Leaves
Leaves opposite or rarely in 3’s; blade narrowly elliptic to broadly ovate or obovate, up to 21 x 12.5 cm, acute or obtuse at the apex, with silvery or yellowish scales; petiole up to c. 5 mm long
Morphology Reproductive morphology Flowers
Flowers 4-merous, in 3–10 cm long axillary spikes
Morphology Reproductive morphology Flowers Receptacle
Lower receptacle 1–3 mm long, tomentose, upper 1.5–2.5 mm long, campanulate, tomentose
Morphology Reproductive morphology Flowers Corolla
Petals 0.2–1.5 mm long, obovate, yellow or greenish, ciliate; filaments 4–6 mm long
Morphology Reproductive morphology Fruits
Fruit 4-winged, 0.8–2.8 cm long, elliptic to subcircular in outline, glabrous to tomentose, with silvery scales; wings papery, 3–12 mm wide; stipe 2–5 mm long.
Distribution
N1–3; C2; S1 widespread in tropical and southern Africa and to Yemen
Ecology
Altitude range 470–1800 m.
Vernacular
Obol (Somali-N).
Note
This is elsewhere in Africa an extremely variable and aggregate species. In Somalia it is not very common and represented by mostly comparatively small-leaved and small-fruited forms.

[FWTA]

Combretaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:1. 1954

Morphology General Habit
Savannah tree with relatively straight bole, to 30 ft. (rarely to 50 ft.) high
Morphology General Bark
Characteristic dark grey deeply fissured bark
Morphology Reproductive morphology Flowers Corolla
Petals and stamens cream.

Native to:

Angola, Benin, Botswana, Burkina, Burundi, Cameroon, Central African Repu, Chad, Congo, Djibouti, Eritrea, Ethiopia, Free State, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Kenya, KwaZulu-Natal, Malawi, Mali, Mozambique, Niger, Nigeria, Northern Provinces, Rwanda, Saudi Arabia, Senegal, Sierra Leone, Somalia, Sudan, Swaziland, Tanzania, Togo, Uganda, Yemen, Zambia, Zaïre, Zimbabwe

Combretum molle R.Br. ex G.Don appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Jun 30, 2010 Sibindo, A.S. [5], Mozambique K000546044
Nov 1, 2008 Cheek, M. [13490], Cameroon K000580112
May 1, 2008 Gilbert [137], Ethiopia K000565475
May 1, 2008 Brummitt [18957], Kenya K000565476
Oct 1, 2003 Onana, J.-M. [1913], Cameroon K000746342
Apr 1, 2002 Pollard, B.J. [441], Cameroon K000746343
Aug 1, 1994 Stuhlmann [774], Tanzania K000226502 lectotype
Jan 1, 1992 Sim, T.R. [6068], Mozambique K000226526 isotype
Jan 1, 1985 Schimper [582], Ethiopia K000226347 Unknown type material
Jan 1, 1980 Kersting [485], Togo K000232571 syntype
Nov 1, 1971 Kotschy [381], Ethiopia K000226464 Unknown type material
Nov 1, 1971 Kotschy [381], Ethiopia K000226465
Apr 1, 1969 Hildebrandt, J.M. [2362], Kenya K000226503 Unknown type material
Mar 1, 1969 Schimper [1358], Ethiopia K000226344 isotype
Mar 1, 1969 Schimper [1358], Ethiopia K000226345 isotype
Mar 1, 1969 Schimper [1358], Ethiopia K000226346 isotype
Feb 1, 1969 Holst, C. [2539], Tanzania K000226500 lectotype
Jan 1, 1967 Miller, O.B. [510], Botswana K000489110
Jan 1, 1967 Reyneke, J. [414], Botswana K000489114
Jan 1, 1961 Welwitsch [4376], Angola K000226517 Unknown type material
Jan 1, 1961 Welwitsch [4315], Angola K000226518
Jan 1, 1961 Welwitsch [4316], Angola K000226519
Jan 1, 1961 Welwitsch [4318], Angola K000226520
Jan 1, 1961 Welwitsch [4312], Angola K000226521
Jan 1, 1961 Baum, H. [155], Angola K000226522 Unknown type material
Jan 1, 1961 Baum, H. [127], Angola K000226523 Unknown type material
Jan 1, 1961 Baum, H. [947], Angola K000226524 Unknown type material
Brenan, J.P.M. [8053], Zambia 25009.000
Brenan, J.P.M. [7977], Zambia 25184.000
Prevett, P.F. [12], Nigeria 3720.000
Schimper [622], Ethiopia K000226339
K000226340 Unknown type material
Schimper [622], Ethiopia K000226341 Unknown type material
Schimper [622], Ethiopia K000226342
Schimper [767], Ethiopia K000226343
Schimper [582], Ethiopia K000226348
Robecchi Bricchetti [25], Somalia K000226466 Unknown type material
Holst, C. [8855], Tanzania K000226497 lectotype
Holst, C. [2916], Tanzania K000226499 Unknown type material
Buchanan, J., C.M.G. [859], Malawi K000226525 lectotype
Gueinzius [567], South Africa K000226651 Unknown type material
Zeyher [575] K000226652 Unknown type material
Burke [521], South Africa K000226653 Unknown type material
Galpin, E.E. [1112], South Africa K000226654 Unknown type material
Timberlake, J. [4936], Mozambique K000545286
Mott, P.J. [230], Botswana K000489109
Hansen, O.J. [3187], Botswana K000489111
Holst, C. [2165], Tanzania K000226498 lectotype
de Beer, J.S. [674], Botswana K000489112
Lea, A. [s.n.], Botswana K000489113
Brunt, M.A. [988], Cameroon K000050782
Schimper [582], Ethiopia K000226349
Brunt, M.A. [878], Cameroon K000050781
Hildebrandt [840], Somalia Combretum somalense K000226467 Unknown type material
Volkens, G. [1662], Tanzania Combretum gueinzii K000226501

First published in Trans. Linn. Soc. London 15: 431 (1827)

Accepted by

  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
  • Boudet, G., Lebrun, J.P. & Demange, R. (1986). Catalogue des plantes vasculaires du Mali: 1-465. Etudes d'Elevage et de Médecine Vétérinaire des Pays Tropicaux.
  • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 3: 1-89. ORSTROM, Bangui.
  • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
  • Edwards, S., Tadesse, M. & Hedberg, I. (eds.) (1995). Flora of Ethiopia and Eritrea 2(2): 1-456. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
  • Hutchinson, J., Dalziel, J.M. & Keay, R.W.J. (1954-1958). Flora of West Tropical Africa, ed. 2, 1: 1-828.
  • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
  • Launert, E. (ed.) (1978). Flora Zambesiaca 4: 1-658. Royal Botanic Gardens, Kew.
  • Liben, L. (1968). Flore du Congo du Rwanda et du Burundi, Combretaceae: 1-105. Jardin Botanique National de Belgique, Bruxelles.
  • Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
  • Wickens, G.E. (1973). Flora of Tropical East Africa, Combretaceae: 1-99.
  • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.

Literature

Flora of West Tropical Africa

  • in Trans. Linn. Soc. 15: 431 (1827).

Kew Backbone Distributions

  • Audru, J., Cesar, J. & Lebrun, J.-P. (1994). Les Plantes Vasculaires de la République de Djibouti. Flore Illustrée 1: 1-336. CIRAD, Départerment d'Elevage et de Médecine vétérinaire, Djibouti.
  • Edwards, S., Tadesse, M. & Hedberg, I. (eds.) (1995). Flora of Ethiopia and Eritrea 2(2): 1-456. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
  • Germishuizen, G. & Meyer, N.L. (eds.) (2003). Plants of Southern Africa an annotated checklist Strelitzia 14: 1-1231. National Botanical Institute, Pretoria.
  • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.

Flora of Somalia

  • Flora Somalia, Vol 1, (1993) Author: by M. Thulin [updated by M. Thulin 2008]

Flora of Tropical East Africa

  • Dale & Greenway, Kenya Trees and Shrubs p. 145 (1961).
  • Exell in Kirkia 7: 189 (1970).
  • F. W. Andr., The Flowering Plants of the Anglo-Egyptian Sudan: 201 (1950).
  • G. Don in Trans. Linn. Soc. 15: 431 (1827)
  • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 137 (1949).
  • Liben in Flore du Congo Belge, du Ruanda et du Burundi, Combr.: 67 (1968).
  • R.Br, in Salt, Voy. Abyss. App.: lxiv (1814), nom. nud.
  • W.J. Eggeling, Indigenous Trees of the Uganda Protectorate, ed. 2: 88 (1952).

Flora Zambesiaca
Flora Zambesiaca
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Flora of Somalia
Flora of Somalia
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of West Tropical Africa
Flora of West Tropical Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0