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This species is accepted, and its native range is South Sudan to Mozambique.
A specimen from Kew's Herbarium

[FZ]

Combretaceae, A. W. Exell. Flora Zambesiaca 4. 1978

Morphology General Habit
Small tree or shrub, sometimes scandent; branchlets densely pubescent when young, soon glabrescent, often silvery-lepidote when young.
Morphology Leaves
Leaves opposite; lamina 3·5–16 x 2–6·5 cm., chartaceous to subcoriaceous, elliptic to elliptic-oblong, minutely verrucose and usually shiny and glabrous above, minutely silvery-lepidote otherwise glabrous beneath, apex usually acuminate, base cuneate or obtuse to rounded; lateral nerves 6–10 pairs, usually slightly impressed above; petiole up to 9(13) mm. long, glabrescent.
Morphology Reproductive morphology Inflorescences
Inflorescence up to 20 cm. long, a terminal or axillary panicle of spikes; rhachis densely fuscous-pubescent when young, ± glabrescent; bracts 1–2 mm. long, linear, caducous.
Morphology Reproductive morphology Flowers
Flowers sessile, white, scented.
Morphology Reproductive morphology Flowers Receptacle
Lower receptacle c. 1 mm. long, glabrous or pubescent; upper receptacle 1·2–1·5 x 1·2–1·5 mm., often petaloid, usually not conspicuously lepidote.
Morphology Reproductive morphology Flowers Calyx
Sepals very shallowly triangular or little developed.
Morphology Reproductive morphology Flowers Corolla
Petals 0· x 1·5 mm., subreniform, glabrous.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamen-filaments 3–4 mm. long; anthers 0·5 mm. long; style 4 mm. long, glabrous.
Morphology Reproductive morphology Fruits
Fruit up to 2·8 x 2·3 cm., broadly elliptic in outline, minutely lepidote otherwise usually glabrous, wings 7 mm. broad, chartaceous, apical peg 1–2 mm. long, stipe up to 5 mm. long.
Morphology General Scales
Scales as for the section, frequent in the areoles of the lower surface of the leaf.

[FTEA]

Combretaceae, G. E. Wickens. Flora of Tropical East Africa. 1973

Morphology General Habit
Small tree or shrub, or in East Africa a scandent shrub or liane.
Morphology Leaves
Leaves opposite; lamina chartaceous to subcoriaceous, elliptic to elliptic-oblong, 3–14(–16) cm. long, 2–6(–6.5) cm. wide, usually acuminate, base cuneate or obtuse to rounded, minutely verrucose and usually shiny and glabrous above, minutely silvery lepidote but otherwise glabrous beneath; lateral nerves 6–10 pairs; petiole up to 9(–13) mm. long.
Morphology Reproductive morphology Inflorescences
Inflorescence terminal and axillary panicles of spikes 2–3 cm. long, in the axils of the upper leaves.
Morphology Reproductive morphology Flowers
Flowers (fig. 3/6, p. 14) white, scented.
Morphology Reproductive morphology Flowers Receptacle
Lower receptacle glabrous or pubescent; upper receptacle 1.2–1.5 mm. long and wide, not conspicuously lepidote.
Morphology Reproductive morphology Flowers Calyx
Sepals very shallowly triangular or little developed.
Morphology Reproductive morphology Flowers Corolla
Petals subreniform, 0.9 mm. long, 1.5 mm. wide, glabrous.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamen-filaments 3–4 mm. long; anthers 0.4 mm. long.
Morphology Reproductive morphology Flowers Disc
Disk inconspicuous, glabrous.
Morphology Reproductive morphology Flowers Gynoecium Style
Style 4–4.5 mm. long, sometimes somewhat flattened near apex.
Morphology Reproductive morphology Fruits
Fruit (fig. 4/6, p. 15) broadly elliptic in outline, up to 2.8 cm. long and 2.3 cm. wide, minutely lepidote, otherwise usually glabrous; wings 7 mm. wide; apical peg 1–2 mm. long;stipe ± 5–8 mm. long.
Morphology General Scales
Scales (fig. 1/6, p. 12) of a simple 8-celled type, 40–50(–70) μ in diameter, frequent in the areoles of the lower leaf-surface.
Habitat
Riverine forest and coastal Brachystegia spiciformis woodland; 10–50 m. and 660–1050 m.
Distribution
P T3 U2 U2/U4

[FTEA]

Combretaceae, G. E. Wickens. Flora of Tropical East Africa. 1973

Morphology General Habit
Scandent shrub or liane, the main branches ending in long leafless whiplike shoots.
Morphology Leaves
Leaves opposite or rarely subopposite; lamina coriaceous, broadly ovate, oblong-ovate or obovate, 3–15 cm. long, 2–7.6 cm. wide, acuminate to rounded at apex, base rounded and slightly emarginate to cuneate, shiny and usually glabrous (except for the scales), often drying a dark reddish brown above, minutely silvery lepidote but otherwise glabrous beneath; scales impressed; lateral nerves 6–9 pairs; domatia present in some of the nerve axils; petiole 4–11 mm. long.
Morphology Reproductive morphology Inflorescences
Inflorescences terminal panicles of spikes 3–6 cm. long, in the axils of the upper leaves.
Morphology Reproductive morphology Flowers
Flowers yellowish.
Morphology Reproductive morphology Flowers Receptacle
Lower receptacle glutinous, shortly pubescent; upper receptacle 1.5 mm. long, 1.5 mm. wide, inconspicuously to densely lepidote.
Morphology Reproductive morphology Flowers Calyx
Sepals very shallowly triangular or little developed.
Morphology Reproductive morphology Flowers Corolla
Petals subreniform to circular, 0.8–1 mm. long, 1.2–1.5 mm. wide, glabrous.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamen-filaments 4–5 mm. long; anthers 0.4 mm. long.
Morphology Reproductive morphology Flowers Disc
Disk inconspicuous, glabrous.
Morphology Reproductive morphology Flowers Gynoecium Style
Style 1.5 mm. long.
Morphology Reproductive morphology Fruits
Fruit (fig. 4/5, p. 15) oblong-ovate to oblong-elliptic in outline, 2.0–2.3 cm. long, 1.8–2.0 cm. wide, apex retuse, base shallowly cordate, minutely lepidote, otherwise glabrous; apical peg inconspicuous; stipe 2–2.5 mm. long.
Morphology General Scales
Scales (fig. 1/5, p. 12) 55–80 μ in diameter, simple 8-celled type with few tangential walls.
Habitat
Riverine forest; 900–1140 m.
Distribution
K4 not known elsewhere

Native to:

Burundi, Kenya, Mozambique, Sudan, Tanzania, Uganda, Zaïre

Combretum umbricola Engl. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
May 1, 1990 Battiscombe, E. [237], Kenya K000226421
May 1, 1990 Battiscombe, E. [237], Kenya K000226422
Jun 1, 1893 Holst, C. [2965], Tanzania K000226420 Unknown type material

First published in Pflanzenw. Ost-Afrikas, C: 288 (1895)

Accepted by

  • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
  • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
  • Jongkind, C.C.H. (2014). Notes on African Combretum Loefl. Species (Combretaceae) Adansonia, sér. 3, 36: 315-327.
  • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
  • Launert, E. (ed.) (1978). Flora Zambesiaca 4: 1-658. Royal Botanic Gardens, Kew.
  • Liben, L. (1968). Flore du Congo du Rwanda et du Burundi, Combretaceae: 1-105. Jardin Botanique National de Belgique, Bruxelles.
  • Wickens, G.E. (1973). Flora of Tropical East Africa, Combretaceae: 1-99.

Literature

Kew Backbone Distributions

  • Jongkind, C.C.H. (2014). Notes on African Combretum Loefl. Species (Combretaceae) Adansonia, sér. 3, 36: 315-327.
  • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.

Flora of Tropical East Africa

  • Burtt Davy, Check-lists Brit. Emp. 1, Uganda: 36 (1935).
  • Engl. in Monographieen Afrikanischer Pflanzen-Familien und Gattungen 3: 23, t. 5/D (1899).
  • Engl., Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete, Theile C: 288 (1895)
  • Exell in Kirkia 7: 173 (1970).
  • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 138 (1949).
  • Liben in Flore du Congo Belge, du Ruanda et du Burundi, Combr.: 56 (1968).

Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0

Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0