- Family: Verbenaceae J.St.-Hil.
- Genus: Phyla Lour.
Phyla nodiflora (L.) Greene
- Genus: Phyla Lour.
[FTEA]
Verbenaceae, B. Verdcourt. Flora of Tropical East Africa. 1992
- Morphology General Habit
- Prostrate creeping herb, with slender often purplish stems 20–45 cm. long covered with fine medifixed hairs; rootstock woody.
- Morphology Leaves
- Leaves often purplish, oblanceolate to obovate, 0.5–5 cm. long, 0.2–3 cm. wide, acute and sharply serrate at the apex, cuneate and entire at the base, glabrous or thinly pubescent with adpressed medifixed hairs.
- Morphology Reproductive morphology Inflorescences
- Spikes purplish, cylindrical, 0.3–2.5 cm. long; bracts broadly obovate to oblate-cuspidate, 2–3 × 3–5 mm.; peduncles 1.3–8 cm. long.
- Morphology Reproductive morphology Flowers Calyx
- Calyx dorsiventrally flattened, 1.4 mm. long, split almost to base abaxially and to about the middle adaxially.
- Morphology Reproductive morphology Flowers Corolla
- Corolla mauve, pink or white with yellow centre, often white and purple in one inflorescence, and sometimes purple in bud, white when open; tube ± 1.5 mm. long; lobes unequal, 0.6–0.8 mm. in diameter.
- Morphology Reproductive morphology Flowers Androecium Stamens
- Stamens 4, didynamous.
- Morphology Reproductive morphology Flowers Gynoecium Stigma
- Stigma obliquely capitate.
- Morphology Reproductive morphology Fruits
- Fruit rounded obovoid, 1.2 × 1–1.2 mm., divided into 2 pyrenes.
- Figures
- Fig. 4.
- Habitat
- Sandy lake shores, shore mud, lava boulders by lakes, lakeside grassland, also roadsides and Avicennia salt marshes; 0–1170 m.
- Distribution
- K3 K5 K7 T2 T3 T4 T5 T6 T8 U1 U2 Z now widespread in tropics, subtropics and warmer temperate regionsthroughout tropical Africa from Senegal, Central African Republic, Sudan, Ethiopia and Somalia to South Africa and also in North Africa.
[CPLC]
Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co
- Distribution
- Nativa en Colombia; Alt. 0 - 363 m.; Islas Caribeñas, Llanura del Caribe, Pacífico, Valle del Cauca, Valle del Magdalena.
- Morphology General Habit
- Hierba
- Conservation
- Preocupación Menor
[FZ]
Flora Zambesiaca. Vol 8, Pt 7. Avicenniaceae, R. Fernandes. Nesogenaceae, M.A. Diniz. Verbenaceae, R. Fernandes. Lamiaceae, R. Fernandes. 2005.
- Type
- Type from N America (Virginia).
- Ecology
- Lake and dam shores, river banks, floodplains, pan margins, coastal dunes and swamp margins, in damp or seasonally inundated soils, sands and heavy black clays and in mud; 0–1300 m.
- Note
- Moldenke, in Fifth Summ. Verbenaceae: 252 (1971), has recorded Phyla nodiflora var. reptans (H.B. & K.) Moldenke for Moçambique. However, we have not been able to confirm this record.
- Distribution
- Widespread in the tropics, subtropics and warmer temperate regions; throughout tropical Africa from Senegal, Central African Republic, Sudan, Ethiopia and Somalia to South Africa. Caprivi Strip
- Morphology General Habit
- Prostrate perennial herb with long trailing stems radiating from a woody taproot. Prostrate perennial herb with long trailing stems radiating from a woody taproot
- Morphology Stem
- Stems up to 2(3) m long, decumbent, terete and somewhat fleshy, drying ± angular and sulcate, rooting at the nodes; internodes 0.7–9 cm long; nodes somewhat thickened; branches 1–2 at each node, from leaf axils, up to 30 cm long, erect or the longer branches ± decumbent and sometimes rooting at the nodes. Stems up to 2(3) m long, decumbent, terete and somewhat fleshy, drying ± angular and sulcate, rooting at the nodes; internodes 0.7–9 cm long; nodes somewhat thickened; branches 1–2 at each node, from leaf axils, up to 30 cm long, erect or the longer branches ± decumbent and sometimes rooting at the nodes
- Morphology General Indumentum
- Indumentum of short appressed medifixed sharply pointed white hairs, sparse on old parts, ± dense on younger parts, mainly on leaves and floral bracts. Indumentum of short appressed medifixed sharply pointed white hairs, sparse on old parts, ± dense on younger parts, mainly on leaves and floral bracts
- Morphology Leaves
- Leaves somewhat fleshy, opposite, sessile or shortly petiolate, 0.8–7.2 × 0.4–2.5 cm, spathulate, oblanceolate or obovate, obtuse to rounded or subacute at the apex and sharply coarsely serrate at the upper part, cuneate and entire at the lower part; petiole up to 8 mm long. Leaves somewhat fleshy, opposite, sessile or shortly petiolate, 0.8–7.2 × 0.4–2.5 cm, spathulate, oblanceolate or obovate, obtuse to rounded or subacute at the apex and sharply coarsely serrate at the upper part, cuneate and entire at the lower part; petiole up to 8 mm long
- Morphology Reproductive morphology Inflorescences
- Spikes short, dense, subglobose or shortly-ovoid at flowering time, and up to 1–2.5(3) cm × 5–9 mm at maturity, cylindric; peduncles (0.5)1–11.5 cm long; bracts closely imbricate, up to c. 2.5 × 2 mm, broadly obovate to oblate-cuspidate, persistent and ± accrescent. Spikes short, dense, subglobose or shortly-ovoid at flowering time, and up to 1–2.5(3) cm × 5–9 mm at maturity, cylindric; peduncles (0.5)1–11.5 cm long; bracts closely imbricate, up to c. 2.5 × 2 mm, broadly obovate to oblate-cuspidate, persistent and ± accrescent
- Morphology Reproductive morphology Flowers Calyx
- Calyx compressed, membranous, deeply split almost to the base abaxially and to about the middle adaxially, persistent but hardly accrescent; lobes 1.5–2 mm long, hyaline or dark purple, shortly toothed on the keels. Calyx compressed, membranous, deeply split almost to the base abaxially and to about the middle adaxially, persistent but hardly accrescent; lobes 1.5–2 mm long, hyaline or dark purple, shortly toothed on the keels
- Morphology Reproductive morphology Flowers Corolla
- Corolla mauve-pink or white with a yellow throat, often white and purple in the same inflorescence, sometimes purple in bud, white when open, c. 2.5 mm long; tube 1.5–2 mm long and 1–1.5 mm broad at the mouth; limb 2–2.5 mm in diameter with the upper lip 2-lobed and erect, the lower lip larger, 3-lobed with the median lobe oblong. Corolla mauve-pink or white with a yellow throat, often white and purple in the same inflorescence, sometimes purple in bud, white when open, c. 2.5 mm long; tube 1.5–2 mm long and 1–1.5 mm broad at the mouth; limb 2–2.5 mm in diameter with the upper lip 2-lobed and erect, the lower lip larger, 3-lobed with the median lobe oblong
- Morphology Reproductive morphology Fruits
- Mericarps 1.5–2 mm long, sub-hemispherical, flattened at the commissural face, obtuse or rounded at the apex, glabrous, enclosed within the persistent calyx. Mericarps 1.5–2 mm long, sub-hemispherical, flattened at the commissural face, obtuse or rounded at the apex, glabrous, enclosed within the persistent calyx.
[FWTA]
Verbenaceae, H. Huber, F. N. Hepper & R. D. Meikle. Flora of West Tropical Africa 2. 1963
- Morphology General Habit
- A prostrate creeping herb
- Ecology
- Of moist places
- Morphology Leaves
- Leaves often tinged purple
- Morphology Reproductive morphology Inflorescences Bracts
- Bracts often tinged purple
- Morphology Reproductive morphology Flowers
- Flowers mauve or pink.
[FSOM]
M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS
- Morphology Stem
- Stems 10–50 cm long, covered by fine appressed hairs
- Morphology Leaves
- Leaves oblanceolate to obovate, 0.5–5 x 0.2–3 cm, serrate in apical part, cuneate and entire in basal part, glabrous or appressed pubescent
- Morphology Reproductive morphology Inflorescences
- Spikes purplish, cylindrical, 0.3–2.5 cm long; bracts broadly obovate to oblate-cuspidate, 2–3 x 3–5 mm; peduncles 0.5–8 cm long
- Morphology Reproductive morphology Flowers Calyx
- Calyx c. 1.4 mm long
- Morphology Reproductive morphology Flowers Corolla
- Corolla mauve, pink or white with yellow centre; tube c. 1.5 mm long; lobes 0.6–0.8 mm in diam. Fruit rounded obovoid, 1.2 x 1–1.2 mm.
- Distribution
- N1–3; C2; S1–3; throughout tropical Africa and also in many other tropical, subtropical and warm temperate areas.
- Ecology
- Altitude range 0–1800 m.
Native to:
Alabama, Albania, Aldabra, Algeria, Andaman Is., Angola, Argentina Northeast, Argentina Northwest, Argentina South, Arizona, Arkansas, Aruba, Assam, Bahamas, Baleares, Bangladesh, Belize, Benin, Bermuda, Bolivia, Botswana, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Burkina, Burundi, California, Cambodia, Cape Provinces, Caprivi Strip, Caroline Is., Cayman Is., Central African Repu, Chad, Chagos Archipelago, Chile Central, Chile North, China South-Central, China Southeast, Colombia, Corse, Costa Rica, Cuba, Cyprus, Djibouti, Dominican Republic, East Himalaya, Ecuador, Egypt, El Salvador, Eritrea, Ethiopia, Florida, Free State, Galápagos, Georgia, Ghana, Greece, Guatemala, Guinea-Bissau, Gulf States, Hainan, Haiti, Honduras, India, Iran, Italy, Jamaica, Japan, Kansas, Kentucky, Kenya, Kriti, KwaZulu-Natal, Laccadive Is., Lebanon-Syria, Leeward Is., Lesotho, Lesser Sunda Is., Liberia, Libya, Louisiana, Madagascar, Malawi, Malaya, Maldives, Mali, Marianas, Mauritania, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Mississippi, Missouri, Morocco, Mozambique, Myanmar, Namibia, Nansei-shoto, Nepal, Netherlands Antilles, Nevada, New Guinea, New Mexico, New South Wales, Nicaragua, Nicobar Is., Niger, Nigeria, North Carolina, Northern Provinces, Northern Territory, Oklahoma, Oman, Oregon, Pakistan, Palestine, Panamá, Paraguay, Pennsylvania, Peru, Philippines, Puerto Rico, Queensland, Samoa, Saudi Arabia, Senegal, Seychelles, Sicilia, Sinai, Socotra, Solomon Is., Somalia, South Carolina, South China Sea, Southwest Caribbean, Spain, Sri Lanka, Sudan, Sulawesi, Swaziland, Tanzania, Texas, Thailand, Tibet, Togo, Trinidad-Tobago, Tunisia, Turkey-in-Europe, Turks-Caicos Is., Uganda, Uruguay, Utah, Venezuela, Venezuelan Antilles, West Himalaya, Western Australia, Windward Is., Zambia, Zaïre, Zimbabwe
Introduced into:
Canary Is., Cocos (Keeling) Is., East Aegean Is., France, Great Britain, Hawaii, Mauritius, New Caledonia, Portugal, Sardegna, South Australia, St.Helena, Victoria
- English
- Daisy lawn, Fog grass
Phyla nodiflora (L.) Greene appears in other Kew resources:
| Date | Reference | Identified As | Barcode | Type Status |
|---|---|---|---|---|
| Meikle, R.D. [1361], Nigeria | 11719.000 | |||
| Bourgeau, E. [1641], Spain | K000215598 | |||
| Reverchon, E. [440], Spain | K000215599 | |||
| Ward, C.J. [44], Angola | K001009799 | |||
| Welwitsch [5638], Angola | K001009806 | |||
| Pearson, H.H.W. [2177], Angola | K001009807 | |||
| Baum, H. [582], Angola | K001009800 | |||
| Pearson, H.H.W. [2177], Angola | K001009808 | |||
| Gossweiler, J. [s.n.], Angola | K001009802 | |||
| Tackholm, V. [8977], Jordan | K000318313 | |||
| Menezes, A. [3794], Angola | K001009804 | |||
| Curror [s.n.], Angola | K001009803 | |||
| Welwitsch [5694], Angola | K001009805 | |||
| Gossweiler, J. [s.n.], Angola | K001009801 | |||
| s.coll. [Cat. no. 1824] | Zappania nodiflora | K001114305 | ||
| s.coll. [Cat. no. 1824] | Zappania nodiflora | K001114307 | ||
| s.coll. [Cat. no. 1824] | Zappania nodiflora | K001114306 | ||
| Wallich, N. [Cat. no. 1824], India | Zappania nodiflora | K001114304 | ||
| s.coll. [Cat. no. 1824] | Zappania nodiflora | K001114308 | ||
| s.coll. [Cat. no. 1824], India | Zappania nodiflora | K001114310 | ||
| s.coll. [Cat. no. 1824] | Zappania nodiflora | K001114309 | ||
| Mar 1, 2013 | Glaziou, A.F.M. [13065], Brazil | Lippia nodiflora | K000925067 | |
| Sellow [s.n.], Brazil | Lippia nodiflora | K000925066 | ||
| Ribas, O.S. [1846], Brazil | Lippia nodiflora | K000925069 | ||
| Sello [2420], Brazil | Lippia nodiflora | K000925070 | ||
| Hatschbach, G.G. [50069], Brazil | Lippia nodiflora | K000925065 | ||
| Sellow, F. [1451], Brazil | Lippia nodiflora | K000925068 | ||
| Glaziou, A.F.M. [13066], Brazil | Lippia nodiflora | K000925064 |
First published in Pittonia 4: 46 (1899)
Accepted by
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- Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
- Cirilo, N. & Proctor, G.R. (1994). Vascular plants of the Caribbean Swan islands of Honduras Brenesia 41-42: 73-80.
- Cossu, T.A, Camarda, I. & Brundu, G. (2014). A catalogue of non-native weeds in irrigated crops in Sardinia (Italy) Webbia; Raccolta de Scritti Botanici 69: 145-156.
- Danin, A. & Fragman- Sapir, O. (2019). Flora of Israel Online http://flora.org.il/en/plants/.
- Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
- Davidse, G. & al. (eds.) (2012). Flora Mesoamericana 4(2): 1-533. Universidad Nacional Autónoma de México, México, D.F.
- Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A., & Tzanoudakis, D. (2013). Vascular plants of Greece. An annotated checklist: 1-372. Botanic gardens and botanical museum Berlin-Dahlem, Berlin and Hellenic botanical society, Athens.
- Fernandes, R. & Diniz, M.A. (2005). Avicenniaceae, Nesogenaceae, Verbenaceae and Lamiaceae (subfams, Viticoideae and Ajugoideae) Flora Zambesiaca 8(7): 1-161. Royal Botanic Gardens, Kew.
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- Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
- Hokche, O., Berry, P.E. & Huber, O. (eds.) (2008). Nuevo Catálogo de la Flora Vascular de Venezuela: 1-859. Fundación Instituto Botánico de Venezuela.
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- MacKee, H.S. (1994). Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie, ed. 2: 1-164. Museum national d'histoire naturelle, Paris.
- Miller, A.G. & Morris, M. (2004). Ethnoflora of Soqotra Archipelago: 1-759. The Royal Botanic Garden, Edinburgh.
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- O'Leary, N. & Múlgura, M.E. (2011). A taxonomic revision of the genus Phyla (Verbenaceae) Annals of the Missouri Botanical Garden 98: 578-596.
- Pandey, R.P. (2009). Floristic diversity of Ferrargunj forest area in South Andaman Journal of Economic and Taxonomic Botany 33: 747-768.
- Raffaelli, M., Mosti, S. & Tardelli, M. (2006). Boswellia sacra Flueck. (Burseraceae) in the Hasik area (Eastern Dhofar, Oman) and a list of surrounding flora Webbia; Raccolta de Scritti Botanici 61: 245-251.
- Shaheen, H., Qureshi, R., Akram, A., Gulfraz, M. & Potter, D. (2014). A preliminary floristic checklist of Thal desert Punjab, Pakistan Pakistan Journal of Botany 46: 13-18.
- Thulin, M. (ed.) (2006). Flora of Somalia 3: 1-626. The Royal Botanic Gardens, Kew.
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Kew Backbone Distributions
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- O'Leary, N. & Múlgura, M.E. (2011). A taxonomic revision of the genus Phyla (Verbenaceae) Annals of the Missouri Botanical Garden 98: 578-596.
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Catálogo de Plantas y Líquenes de Colombia
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