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This genus is accepted, and its native range is Cosmopolitan.
Polystichum craspedosorum


Dryopteridaceae, JP Roux, M Shaffer-Fehre, B Verdcourt. Flora of Tropical East Africa. 2007

Morphology General Habit
Terrestrial or lithophytic ferns, rarely epiphytic
Vegetative Multiplication Rhizomes
Rhizome erect or creeping, with dense scales
Morphology Leaves
Fronds tufted or spaced.
Morphology Leaves Stipes
Stipe scaly when young, glabrescent, densely covered in scales.
Morphology Leaves Leaf lamina
Lamina pinnate to 3-pinnate, often proliferous near apex.
Morphology Leaves Pinnae
Pinnae opposite to alternate, petiolulate.
Morphology Leaves Rachis
Rachis densely covered in scales; venation free, anadromous
Morphology Reproductive morphology Sori
Sori borne medially near or at vein ending, circular, indusium peltate or rarely absent.
160–200 species throughout temperate zone and in tropical mountains.

Native to:

Afghanistan, Alabama, Alaska, Albania, Alberta, Aleutian Is., Algeria, Altay, Amsterdam-St.Paul Is, Antipodean Is., Argentina Northeast, Argentina Northwest, Argentina South, Arizona, Arkansas, Assam, Austria, Azores, Baleares, Baltic States, Belarus, Belgium, Bismarck Archipelago, Bolivia, Borneo, Brazil South, Brazil Southeast, Brazil West-Central, British Columbia, Bulgaria, Buryatiya, California, Cameroon, Canary Is., Cape Provinces, Central European Rus, Chatham Is., Chile Central, Chile South, China North-Central, China South-Central, China Southeast, Colombia, Colorado, Comoros, Connecticut, Corse, Costa Rica, Crozet Is., Cuba, Czechoslovakia, Delaware, Denmark, District of Columbia, Dominican Republic, East European Russia, East Himalaya, Easter Is., Ecuador, El Salvador, Ethiopia, Falkland Is., Fiji, Finland, Florida, France, Free State, Føroyar, Galápagos, Georgia, Germany, Great Britain, Greece, Greenland, Guatemala, Gulf of Guinea Is., Hainan, Haiti, Hawaii, Honduras, Hungary, Iceland, Idaho, Illinois, India, Indiana, Inner Mongolia, Iowa, Iran, Iraq, Ireland, Irkutsk, Italy, Jamaica, Japan, Jawa, Juan Fernández Is., Kamchatka, Kansas, Kazakhstan, Kentucky, Kenya, Khabarovsk, Korea, Krasnoyarsk, Kriti, Krym, Kuril Is., KwaZulu-Natal, Labrador, Lebanon-Syria, Leeward Is., Lesotho, Lesser Sunda Is., Louisiana, Macquarie Is., Madagascar, Madeira, Maine, Malawi, Malaya, Maluku, Manchuria, Marion-Prince Edward, Marquesas, Maryland, Massachusetts, Mauritius, Mexican Pacific Is., Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Northwest, Mexico Southeast, Mexico Southwest, Michigan, Minnesota, Mississippi, Missouri, Montana, Morocco, Mozambique, Myanmar, Nansei-shoto, Nepal, Netherlands, Nevada, New Brunswick, New Caledonia, New Guinea, New Hampshire, New Jersey, New South Wales, New York, New Zealand North, New Zealand South, Newfoundland, Nicaragua, Norfolk Is., North Carolina, North Caucasus, North European Russi, Northern Provinces, Northwest European R, Northwest Territorie, Norway, Nova Scotia, Ohio, Oklahoma, Ontario, Oregon, Pakistan, Panamá, Paraguay, Pennsylvania, Peru, Philippines, Poland, Portugal, Primorye, Prince Edward I., Puerto Rico, Qinghai, Queensland, Québec, Rhode I., Romania, Rwanda, Réunion, Sakhalin, Sardegna, Sicilia, Society Is., Solomon Is., Somalia, South Carolina, South Dakota, South Georgia, Spain, Sri Lanka, Sudan, Sulawesi, Sumatera, Swaziland, Sweden, Switzerland, Tadzhikistan, Taiwan, Tanzania, Tasmania, Tennessee, Texas, Thailand, Tibet, Transcaucasus, Tristan da Cunha, Tubuai Is., Tunisia, Turkey, Turkey-in-Europe, Uganda, Ukraine, Uruguay, Utah, Venezuela, Vermont, Victoria, Vietnam, Virginia, Washington, West Himalaya, West Siberia, West Virginia, Wisconsin, Wyoming, Xinjiang, Yemen, Yugoslavia, Yukon, Zambia, Zaïre, Zimbabwe

Polystichum Roth appears in other Kew resources:

First published in Tent. Fl. Germ. 3(1): 31, 69 (1799)

Accepted by

  • Fraser-Jenkins, C.R. & Kandel, D.R. (2019). Ferns and Fern-Allies of Nepal 2: 1-446. Government of Nepal, Ministry of Forests and Environment, Department of Plant Resources.
  • Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.


Flora of Tropical East Africa

  • Roux, Polystichum in Africa: 1–375 (1998)
  • Tent. Fl. Germ. 3: 31 (1799);

Flora of Tropical East Africa
Flora of Tropical East Africa

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 World Checklist of Selected Plant Families.

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.