- Family:
- Phyllanthaceae Martinov
Breynia J.R.Forst. & G.Forst.

[FZ]
Euphorbiaceae, A. Radcliffe-Smith. Flora Zambesiaca 9:4. 1996
- Morphology Leaves
- Leaves alternate, shortly petiolate, stipulate, simple, entire, penninerved, borne on plagiotropic shoots (leafy or floriferous lateral shoots of limited growth, see Phyllanthus).
- Morphology Reproductive morphology Flowers
- Flowers axillary, the male fasciculate or solitary, usually in the proximal axils, the female solitary, usually in the distal axils. Male flowers: pedicels often capillary; calyx obconic or turbinate, calyx lobes 6, imbricate, sharply inflexed; petals absent; disk absent; stamens 3, united into a short column, anthers elongate, thecae linear, extrorse, adnate to the column, longitudinally dehiscent; pistillode absent. Female flowers: pedicels sometimes capillary; calyx lobes 6, imbricate, not inflexed, usually larger than in the male flowers, accrescent; disk absent; ovary 3-locular, ovules 2 per locule; styles 3, free, short, erect, simple or bifid.
- Morphology Reproductive morphology Fruits
- Fruit ± baccate, tardily and often incompletely loculicidally dehiscent; exocarp sometimes somewhat fleshy; endocarp crustaceous.
- Morphology Reproductive morphology Seeds
- Seeds trigonous, ecarunculate; testa membranous; albumen fleshy; cotyledons broad; radicle long.
- Morphology General Habit
- Monoecious or apparently dioecious shrubs or small trees, with or without a simple indumentum, often blackening on drying.
- Morphology Stem
- Branching phyllanthoid (see Tab. 8).
[FTEA]
Euphorbiaceae, A. R.-Smith. Flora of Tropical East Africa. 1987
- Morphology General Habit
- Monoecious or apparently dioecious shrubs or small trees with or without a simple indumentum, often blackening on drying
- Morphology Leaves
- Cataphylls present, as in Phyllanthus (q.v.) Leaves alternate, shortly petiolate, stipulate, simple, entire, penninerved, arranged on shoots of limited growth (plagiotropic)
- Morphology Reproductive morphology Flowers
- Flowers axillary, the males fasciculate or solitary in the proximal axils, the females solitary in the distal axils Female flowers: pedicels capillary or not; calyx-lobes 6, imbricate, not inflexed, usually larger than those of the ♂ flower, accrescent; disc 0; ovary 3-locular, with 2 ovules per locule; styles 3, free, short, erect, simple or bifid Male flowers: pedicels often capillary; calyx obconic or turbinate; calyx-lobes 6, imbricate, the distal halves sharply inflexed with their apices almost touching the androecium; petals 0; disc 0; stamens 3, united to form a short column, anthers elongate, the thecae linear, extrorse, adnate to the column, longitudinally dehiscent; pistillode 0
- sex Male
- Male flowers: pedicels often capillary; calyx obconic or turbinate; calyx-lobes 6, imbricate, the distal halves sharply inflexed with their apices almost touching the androecium; petals 0; disc 0; stamens 3, united to form a short column, anthers elongate, the thecae linear, extrorse, adnate to the column, longitudinally dehiscent; pistillode 0
- sex Female
- Female flowers: pedicels capillary or not; calyx-lobes 6, imbricate, not inflexed, usually larger than those of the ♂ flower, accrescent; disc 0; ovary 3-locular, with 2 ovules per locule; styles 3, free, short, erect, simple or bifid
- Morphology Reproductive morphology Fruits
- Fruit tardily and often incompletely loculicidally dehiscent; exocarp sometimes somewhat fleshy; endocarp crustaceous
- Morphology Reproductive morphology Seeds
- Seeds trigonous, ecarunculate.
Native to:
Andaman Is., Assam, Bangladesh, Bismarck Archipelago, Borneo, Cambodia, China South-Central, China Southeast, East Himalaya, Hainan, India, Jawa, Laos, Lesser Sunda Is., Malaya, Maluku, Myanmar, Nansei-shoto, Nepal, New Caledonia, New Guinea, New South Wales, Nicobar Is., Northern Territory, Pakistan, Philippines, Queensland, Réunion, Solomon Is., Sri Lanka, Sulawesi, Sumatera, Taiwan, Thailand, Tibet, Vanuatu, Vietnam
Introduced into:
Bolivia, Caroline Is., Chagos Archipelago, Christmas I., Cocos (Keeling) Is., Cook Is., Cuba, Dominican Republic, Fiji, Florida, Gambia, Guinea, Gulf of Guinea Is., Haiti, Hawaii, Leeward Is., Line Is., Marianas, Marshall Is., Niue, Norfolk Is., Ogasawara-shoto, Puerto Rico, Samoa, Seychelles, Society Is., Trinidad-Tobago, Tuamotu, Windward Is.
- Breynia amabilis (Airy Shaw) Welzen & Pruesapan
- Breynia amoebiflora (Airy Shaw) Welzen & Pruesapan
- Breynia androgyna (L.) Chakrab. & N.P.Balakr.
- Breynia assimilis (Thwaites) Chakrab. & N.P.Balakr.
- Breynia asteranthos (Airy Shaw) Welzen & Pruesapan
- Breynia asymmetrica (Welzen) Welzen & Pruesapan
- Breynia baudouinii Beille
- Breynia beillei Welzen & Pruesapan
- Breynia bicolor (Craib) Chakrab. & N.P.Balakr.
- Breynia bishnupadae (M.Gangop. & Chakrab.) Chakrab. & N.P.Balakr.
- Breynia bonii (Beille) Welzen & Pruesapan
- Breynia brevipes (Müll.Arg.) Chakrab. & N.P.Balakr.
- Breynia calcarea (M.R.Hend.) Welzen & Pruesapan
- Breynia carnosa Welzen & Pruesapan
- Breynia cernua (Poir.) Müll.Arg.
- Breynia collaris Airy Shaw
- Breynia coriacea Beille
- Breynia coronata Hook.f.
- Breynia delavayi (Croizat) Welzen & Pruesapan
- Breynia discigera Müll.Arg.
- Breynia discocalyx (Welzen) Welzen & Pruesapan
- Breynia disticha J.R.Forst. & G.Forst.
- Breynia diversifolia Beille
- Breynia fleuryi Beille
- Breynia fruticosa (L.) Müll.Arg.
- Breynia garrettii (Craib) Chakrab. & N.P.Balakr.
- Breynia glauca Craib
- Breynia gour-maitii (Chakrab. & M.Gangop.) Chakrab. & N.P.Balakr.
- Breynia grandiflora Beille
- Breynia granulosa (Airy Shaw) Welzen & Pruesapan
- Breynia gynophora Welzen & Pruesapan
- Breynia harmandii (Beille) Welzen & Pruesapan
- Breynia heteroblasta (Airy Shaw) Welzen & Pruesapan
- Breynia heyneana J.J.Sm.
- Breynia hirsuta (Beille) Welzen & Pruesapan
- Breynia indosinensis Beille
- Breynia kerrii (Airy Shaw) Welzen & Pruesapan
- Breynia kitanovii (Thin) Welzen & Pruesapan
- Breynia lanceolata (Hook.f.) Welzen & Pruesapan
- Breynia lithophila Welzen & Pruesapan
- Breynia macrantha (Hassk.) Chakrab. & N.P.Balakr.
- Breynia maichauensis (Thin) Welzen & Pruesapan
- Breynia massiei Beille
- Breynia micrasterias (Airy Shaw) Welzen & Pruesapan
- Breynia microphylla (Kurz ex Teijsm. & Binn.) Müll.Arg.
- Breynia mollis J.J.Sm.
- Breynia oblongifolia (Müll.Arg.) Müll.Arg.
- Breynia obscura Welzen & Pruesapan
- Breynia officinalis Hemsl.
- Breynia orbicularis (Craib) Welzen & Pruesapan
- Breynia pierrei (Beille) Welzen & Pruesapan
- Breynia platycalyx Airy Shaw
- Breynia po-khantii (Chakrab. & M.Gangop.) Chakrab. & N.P.Balakr.
- Breynia podocarpa Airy Shaw
- Breynia poilanei (Beille) Welzen & Pruesapan
- Breynia poomae (Welzen & Chayam.) Welzen & Pruesapan
- Breynia pubescens Merr.
- Breynia pulchella (Airy Shaw) Welzen & Pruesapan
- Breynia quadrangularis (Willd.) Chakrab. & N.P.Balakr.
- Breynia racemosa (Blume) Müll.Arg.
- Breynia repanda (Müll.Arg.) Chakrab. & N.P.Balakr.
- Breynia repens Welzen & Pruesapan
- Breynia reticulata (S.L.Mo ex P.T.Li) Welzen & Pruesapan
- Breynia retroversa (Wight) Chakrab. & N.P.Balakr.
- Breynia retusa (Dennst.) Alston
- Breynia rhynchocarpa Benth.
- Breynia rigida (Thwaites) Chakrab. & N.P.Balakr.
- Breynia rostrata Merr.
- Breynia saksenana (Manilal, Prasann. & Sivar.) Chakrab. & N.P.Balakr.
- Breynia septata Beille
- Breynia shawii (Welzen) Welzen & Pruesapan
- Breynia similis (Craib) Welzen & Pruesapan
- Breynia spatulifolia (Beille) Welzen & Pruesapan
- Breynia stipitata Müll.Arg.
- Breynia subangustifolia Thin
- Breynia suberosa (Airy Shaw) Welzen & Pruesapan
- Breynia subindochinensis Thin
- Breynia subterblanca (C.E.C.Fisch.) C.E.C.Fisch.
- Breynia temii (Welzen & Chayam.) Welzen & Pruesapan
- Breynia thoii (Thin) Welzen & Pruesapan
- Breynia thorelii (Beille) Welzen & Pruesapan
- Breynia thyrsiflora (Welzen) Welzen & Pruesapan
- Breynia tiepii Welzen & Pruesapan
- Breynia tonkinensis Beille
- Breynia trinervia (Hook.f. & Thomson ex Müll.Arg.) Chakrab. & N.P.Balakr.
- Breynia tsiangii (P.T.Li) Welzen & Pruesapan
- Breynia vestita Warb.
- Breynia villosa (Blanco) Welzen & Pruesapan
- Breynia virgata (Blume) Müll.Arg.
- Breynia vitis-idaea (Burm.f.) C.E.C.Fisch.
- Breynia yanhuiana (P.T.Li) Welzen & Pruesapan
Breynia J.R.Forst. & G.Forst. appears in other Kew resources:
Date | Reference | Identified As | Barcode | Type Status | Has image? |
---|---|---|---|---|---|
Johns, R.J. [8916], New Guinea | 70224.000 | No |
First published in Char. Gen. Pl.: 145 (1776)
Accepted by
- Govaerts, R. (1996). World Checklist of Seed Plants 2(1, 2): 1-492. MIM, Deurne.
- Govaerts, R., Frodin, D.G. & Radcliffe-Smith, A. (2000). World Checklist and Bibliography of Euphorbiaceae (and Pandaceae) 1-4: 1-1622. The Board of Trustees of the Royal Botanic Gardens, Kew.
- van Welzen, P.C., Pruesapan, K., Telford, I.R.H., Esser, H.-J. & Bruhl, J.J. (2014). Phylogenetic reconstruction prompts taxonomic changes in Sauropus, Synostemon and Breynia (Phyllanthaceae tribe Phyllantheae). Blumea 59: 77-94.
Literature
Flora Zambesiaca
- Char. Gen. Pl.: 145, t. 73 (1776).
- Müller Argoviensis in De Candolle, Prodr. 15, 2: 438 (1866), nom. conserv.
Flora of Tropical East Africa
- Char. Gen. Pl.: 145, t. 73 (1776)
- G.P. 3(1): 276 (1880), nom. conserv.
- Muell. Arg. in DC., Prodr. 15(2): 438 (1866)
-
Flora Zambesiaca
Flora Zambesiaca
http://creativecommons.org/licenses/by-nc-sa/3.0
-
Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0
-
Herbarium Catalogue Specimens
-
Interactive Key to Seed Plants of Malesia and Indo-China
The Malesian Key Group (2010) Interactive Key to Seed Plants of Malesia and Indo-China (Version 2.0, 28 Jul 2010) The Nationaal Herbarium Nederland Leiden and The Royal Botanic Gardens Kew
-
Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0
-
Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2022. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0