Leucas calostachys Oliv.

Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Leucas, G. Bramley. Flora of Tropical East Africa. 2009

Type

Type: Tanzania, Bukoba District: Karagwe, Grant in Speke & Grant 413 (K!, holo.)

Morphology General Habit

Erect shrub with branching stems, 0.6–2.5 m high, from a woody rootstock

Morphology Stem

Stems tomentose or pubescent with appressed upward pointing or with long patent hairs, sometimes becoming less pubescent as they mature

Morphology Leaves

Leaves petiolate, often shortly so, occasionally or often on short shoots in the leaf axils; blades almost orbicular to elliptic to obovate to ± spathulate, 0.9–6 × 0.6–3.5 cm, margins crenate or crenateserrate towards apex, becoming entire towards base, apex obtuse, base attenuate, upper surface densely pubescent but surface not obscured, lower surface tomentose to velutinous, the surface ± obscured or occasionally densely pubescent, surface not obscured and venation visible, eglandular; petioles 5–7 mm long, tomentose or velutinous

Morphology Reproductive morphology Inflorescences

Inflorescence of 3–16 verticils spaced 3–60 mm apart, forming a dense elongate cluster or laxly spaced; many (>15) flowers per verticil; pedicels 0.5–1.5 mm long, pubescent with appressed hair; bracteoles linear to lanceolate, 3–9 mm long, pubescent with appressed hair

Morphology Reproductive morphology Flowers Calyx

Calyx funnel-shaped, 9–11 mm long (tube 5–7 mm) at anthesis, expanding 1–2 mm in fruit, outer surface velutinous or densely pubescent with long appressed hair when in bud and flower, less conspicuously hairy in fruit, with occasional red sessile glands, inner surface with short hairs; throat asymmetric, anterior lip extending ± 4 mm from the end of the tube; posterior lip with five triangular to narrowly triangular lobes, these shortly subulate at the apex, ± 0.5 mm long, the central lobe slightly larger, two narrowly triangular lateral lobes present between the posterior and anterior lips, ± 0.5 mm long, the anterior lip divided into three triangular lobes, shortly subulate at the apex, ± 1 mm long

Morphology Reproductive morphology Flowers Corolla

Corolla white, 9–13 mm long; tube 5–7 mm long, pubescent with short hair, eglandular, inner surface with patches of glandular hairs leading to an undulate band of glandular hairs halfway down the tube; posterior lip 4–7 mm long; anterior lip 3–6 mm long, inner surface with patches of short glandular hairs, outer surface with longer appressed hairs except on the backs on the lobes

Morphology Reproductive morphology Fruits

Nutlets dark brown, 1.5–2 × 1 mm, gland-dotted with more distinctive glands and a few hairs on the flattened top

Figures

Fig 11: 6, p 55