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This species is accepted, and its native range is Tropical & Subtropical Old World.
Ocimum gratissimum

[FWTA]

Labiatae, J. K. Morton. Flora of West Tropical Africa 2. 1963

Diagnostic
Similar to O. suave but subglabrous
Ecology
Usually occurring around villages.
Vernacular
The Tea Bush.

[FSOM]

M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS

Morphology General Habit
Perennial herb with a woody base, 0.4–2.5 m tall, with simple or rarely also some branched hairs
Morphology Leaves
Leaves petiolate; blade elliptic or ovate to broadly ovate, up to 30–150 × 20–85 mm, apex acute, acuminate or obtuse, base cuneate or attenuate, margin serrate
Morphology Reproductive morphology Inflorescences
Inflorescence lax or dense; bracts becoming downwards-pointing, deciduous after anthesis
Morphology Reproductive morphology Fruits Fruiting calyx
Fruiting calyx mostly ± downwards-pointing, 5–6 mm long, closed by the lower lobes being pressed against the upper lobe; upper lobe widest near the apex, 3–4 mm broad, narrowly cuneate at the base; lateral lobes subulate at apex; lower lobes connate to an emarginate lip
Morphology Reproductive morphology Flowers Corolla
Corolla 3–5 mm long, greenish, dull white or pale yellowish
Morphology Reproductive morphology Flowers Androecium Stamens
Anterior stamens c. 1 mm longer than the lower lip of the corolla
Morphology Reproductive morphology Fruits
Nutlets c. 1.5 × 1.5 mm, glabrous, minutely tuberculate, hardly producing mucilage when wet.
Note
Paton (1992) divided O. gratissimum into two subspecies. The description above refers to subsp. gratissimum, the only subspecies known from Somalia.

[KBu]

Suddee, S., A. J. Paton, & Parnell, J. (2005). Taxonomic Revision of Tribe Ocimeae Dumort. (Lamiaceae) in Continental South East Asia III. Ociminae. Kew Bulletin, 60(1), 3-75. Retrieved from http://www.jstor.org/stable/4110885

Morphology General Habit
Perennial herbs, undershrubs or shrubs up to 2 m tall
Morphology Leaves
Leaves spreading or ascending, ovate or elliptic-ovate, 25 - 100 x 12 - 60 mm, apex acute, base cuneate or attenuate, margin sparsely serrate, glandular-punctate or not, glabrous, pubescent or tomentose on both sides, sometimes glabrous or puberulent above, with scattered hairs restricted to veins beneath; petiole 10 - 50 mm long, slender, glabrous or pubescent
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens with posterior pair having a hairy transverse process near base
Morphology Reproductive morphology Flowers Calyx
Calyx 2 - 3 mm long at anthesis, 3 - 4 mm long in fruit; posterior lip rounded, accrescent, decurrent on tube, apex pointed; anterior lip much shorter than posterior, the 2 lateral teeth acute, level with or below the 2 median teeth which are pressed against the posterior lip; throat closed; tube with short patent hairs with or without sessile glands outside, pubescent inside
Morphology Reproductive morphology Flowers Corolla
Corolla greenish- white, 3 - 4 mm long, lobes obscurely crenate, with hairs on back, with or without sessile glands; posterior lip oblong, lobes equal; anterior lip boat shaped; tube straight, puberulous outside, glabrous inside
Morphology Reproductive morphology Fruits
Nutlets brown, subglobose, minutely tuberculate, producing mucilage when wet.
Morphology Reproductive morphology Inflorescences
Inflorescence lax or dense, verticils up to 10 mm apart, axis glabrous or softly pubescent; bracts caducous, ovate with very broad base, 2 - 5 mm long, apex acuminate, base cuneate or sessile, margin ciliate, pubescent on both sides, with 3 nerves at base; pedicels 2 - 3 mm long, recurved, pubescent
Morphology Stem
Stems usually woody at base, round-quadrangular, glabrous, glabrescent or pubescent
Type
Cultivated in Uppsala, originally from India, Linnean Herbarium 749.2 [neotype LINN, (microfiche!)].

[FTEA]

Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Flora of Tropical East Africa. 2009

Type
Neotype, chosen by Paton, loc. cit. (1992) – specimen probably a later addition to Linnean Herbarium: cultivated in Uppsala, originally from India, Linnean Herbarium 749.2 (LINN!, neo.)
Morphology General Habit
Aromatic perennial herb 0.6–2.5 m tall
Morphology Stem
Stems erect, rounded-quadrangular, much branched, often striate, woody at base with epidermis often peeling in strips; glabrous or with scattered patent hairs below, becoming pubescent at nodes and on the inflorescence-axis, or pubescent with ± retrorse and patent hairs becoming hoary or tomentose on the inflorescence axis
Morphology Leaves
Leaf blades often darker above, elliptic or ovate to broadly ovate, 1.5–15 × 1–8.5 cm, serrate, apex obtuse, acute or acuminate, base cuneate or attenuate, indumentum of scattered short hairs mostly confined to veins beneath, or lower surface pubescent with upper glabrous or pubescent overall, sometimes tomentose beneath; petiole 5–30 mm long, glandular-punctate or not
Morphology Reproductive morphology Inflorescences
Inflorescence lax or dense (calyces of adjacent verticils touching or verticils ± 1 cm apart); bracts erect above forming a small green terminal coma, becoming downward-pointing and deciduous towards base, ovate or narrowly ovate, 3–12 × 1–7 mm, acuminate, sometimes cucullate; pedicels 3–4 mm long, spreading or ascending, slightly curved, ± flattened
Morphology Reproductive morphology Flowers Calyx
Calyx horizontal, ± downward-pointing or strongly reflexed against the inflorescence axis, 2–3 mm long at anthesis, pubescent, hoary or tomentose, with sessile glands or not; posterior lip large, rounded at tip, decurrent, teeth of anterior lip lanceolate, acuminate, teeth of lateral lobes bristle-pointed; fruiting calyx 6–8 mm long, tube closed, posterior lip accrescent, rounded or obtuse and wider at tip, decurrent, median lobes of anterior lip pressed against the posterior lip, teeth of the lateral lobes level with, or lower than, the underside of the anterior lip, sometimes spreading; indumentum pubescent, hoary or tomentose with sessile glands or not
Morphology Reproductive morphology Flowers Corolla
Corolla greeny or dull white or pale yellow, 3–4 mm long; tube straight, funnel-shaped, scarcely exceeding the calyx tube
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens exserted 1 mm from corolla, posterior with a flattened, hairy or subglabrous appendage near base
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary glabrous
Morphology Reproductive morphology Fruits
Nutlets brown, ± spherical, 1.5 mm in diameter, minutely tuberculate, producing a small amount of mucilage when wet
Figures
Fig 22: 7, 8, p 139
Note
(on the species as a whole). O. gratissimum forms a polymorphic complex in Africa. Subsp. iringense is easily recognised by its strongly reflexed fruiting calyces. Within subsp. gratissimum, var. macrophyllum is delimited from var. gratissimum by the combination of a lax inflorescence and sparse indumentum. This delimitation is supported by reference to Indian material where the discontinuity between the two varieties of subsp. gratissimum is upheld and the fact that var. macrophyllum has a limited geographical distribution in East Africa. Var. macrophyllum may have been introduced either from or to India and spread throughout Africa by cultivation. Cultivated Labiatae often tend to be glabrous as a result of selection by man. However, it is also possible that it may have arisen from var. gratissimum several times in response to environmental factors. Thus varietal rank is preferred to subspecific. See also Vieira, R., Grayer, R.J., Paton, A., and Simon, J.E. (2001) – Genetic diversity of Ocimum gratissimum L. based on volatile oil contents, flavonoids and RAPD markers. Biochemical Systematics and Ecology 29: 287–304. Forms exist in Uganda and Tanzania that correspond to var. macrophyllum in leaf and calyx indumentum and in having a lax inflorescence, but with a stem indumentum similar to var. gratissimum. These forms are similar to the type of O. gratissimum. There is no discontinuity that can separate these forms from the more densely hairy to tomentose forms previously referred to as O. urticifolium. Nor is there any morphological or ecological discontinuity separating forms with dense, wide, robust inflorescences (previously referred to as O. trichodon) from specimens with narrower inflorescences ( O. urticifolium). Thus within var. gratissimum inflorescence structure and stem, leaf and calyx indumentum form a continuum of variation. Two varieties based on the types of O. gratissimum and either O. urticifolium or O. trichodon could be recognised as there are large morphological differences between the extremes of this continuum. There are also supporting differences in leaf epidermis structure (Olowokudejo & Pereira-Shateolu in Phytomorphology 38: 147 (1988)) and chromosome number (Morton in J.L.S. 58: 231 (1962)). However, the practical difficulties of dividing and recognising the divisions of this continuum and the interbreeding between members of this complex reported by Khosta in J. Pl. Anat. Morphol. 3: 157 (1986) make this undesirable.

[UNAL]
Vernacular
Albahaca vaca, Atiyayo, Basen, Fonbazen, Vanne van

[CPLC]

Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

Distribution
Cultivada en Colombia; Alt. 300 - 1600 m.; Amazonia, Andes, Pacífico, Valle del Cauca.
Morphology General Habit
Hierba, subarbusto

Native to:

Aldabra, Andaman Is., Angola, Assam, Bangladesh, Botswana, Burundi, Cambodia, Cameroon, Caprivi Strip, Central African Repu, China Southeast, Comoros, Congo, Djibouti, Equatorial Guinea, Eritrea, Ethiopia, Gabon, Ghana, Guinea, Gulf of Guinea Is., India, Ivory Coast, Jawa, Kenya, KwaZulu-Natal, Laccadive Is., Laos, Liberia, Madagascar, Malawi, Malaya, Maldives, Mauritius, Mozambique, Namibia, Nepal, Nigeria, Northern Provinces, Rwanda, Senegal, Sierra Leone, Somalia, Sri Lanka, Sudan, Sumatera, Swaziland, Taiwan, Tanzania, Thailand, Togo, Uganda, Vietnam, Yemen, Zambia, Zaïre, Zimbabwe

Introduced into:

Aruba, Benin, Bismarck Archipelago, Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Burkina, Cape Verde, Chad, Cook Is., Cuba, Dominican Republic, Gambia, Guinea-Bissau, Hawaii, Jamaica, Leeward Is., Marquesas, Mexico Gulf, Mexico Southwest, Netherlands Antilles, New Caledonia, New Guinea, Niger, Panamá, Queensland, Society Is., Tonga, Trinidad-Tobago, Tuamotu, Windward Is.

Ocimum gratissimum L. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Nov 1, 2013 Ende [2784], China K000938238
Nov 1, 2013 David, B.et al. [CL655], Cambodia K000938295
Nov 1, 2013 Maraia [AM050], Madagascar K000938312
Jan 1, 2011 Mwangoka [5224], Brazil K000735148
Jan 1, 2011 Du, N.V. et al. [HNK2612], Vietnam K000735158
Jan 1, 2010 Betz, W. [31], Papua New Guinea K000897217
Aug 15, 2006 Katende [643], Uganda K000194745
May 23, 2006 Folli [3235], Brazil K000194505
Nov 29, 1999 Takeuchi, W. [13813], Papua New Guinea K000897347
Nov 29, 1999 Takeuchi, W. [13813], Papua New Guinea K000897348
Dec 1, 1997 Etuge, M. [3702], Cameroon K000339266
Apr 1, 1964 Haniff, M. [SFN6923], Malaysia K000897352
Waterhouse, J.H.L. [916], Papua New Guinea K000897344
s.coll. [Cat. no. 2720] K001116936
s.coll. [Cat. no. 2720] K001116933
Wiakabu, J. [LAE73476], Papua New Guinea K000897349
s.coll. [Cat. no. 2720], India K001116938
s.coll. [Cat. no. 2720] K001116934
s.coll. [Cat. no. 2720], India K001116937
Stevens, P.F. [LAE50142], Papua New Guinea K000897350
Streimann, H. [NGF27573], Papua New Guinea K000897343
Wallich, N. [Cat. no. 2720], Malaysia K001116930
Kerenga, K. [LAE77444], Papua New Guinea K000897346
s.coll. [Cat. no. 2720] K001116932
Peekel, G. [12], Papua New Guinea K000897342
s.coll. [Cat. no. 2720] K001116931
s.coll. [Cat. no. 2720] K001116935
Coode, M. [NGF29791], Papua New Guinea K000897345

First published in Sp. Pl.: 1197 (1753)

Accepted by

  • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Balkrishna, A. (2018). Flora of Morni Hills (Research & Possibilities): 1-581. Divya Yoga Mandir Trust.
  • Berhaut, J. (1975). Flore illustrée du Sénégal 4: 1-625. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
  • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 2(1): 1-85. ORSTOM, Bangui.
  • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
  • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
  • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
  • Davidse, G. & al. (eds.) (2012). Flora Mesoamericana 4(2): 1-533. Universidad Nacional Autónoma de México, México, D.F.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
  • Garcia-Mendoza, A.J. & Meave, J.A. (eds.) (2012). Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2: 1-351. Instituto de Biología, Universidad Nacional Autónoma de México.
  • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
  • Harris, D.J. (2002). The vascular plants of the Dzanga-Sangha Reserve, Central African Republic: 1-274. National Botanic Garden (Belgium), Meise.
  • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
  • Khanam, M. & Hassan, M.A. (2008). Lamiaceae Flora of Bangladesh 58: 1-161. Bangladesh National Herbarium, Dhaka.
  • Lebrun, J.P., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Mabberley, D.J. & De Kok, R.P.J. (2004). Labiatae Flore de la Nouvelle-Calédonie et Dépendances 25: 20-141. Muséum National d'Histoire Naturelle, Paris.
  • Morales, R. (2011). Les Labiadas (Lamiaceae) de Guinea Ecuatorial Anales del Jardin Botanico de Madrid 68: 199-223.
  • Paton, A.J., Bramley, G., Ryding, O., Polhill, R., Harvey, Y., Iwarsson, M., Willis, F., Phillipson, P., Balkwill, K., Lukhoba, C., Otiend, D & Harley (2009). Lamiaceae (Labiatae) Flora of Tropical East Africa: 1-430.
  • Paton, A.J., Bramley, G., Ryding, O., Polhill, R.M., Harvey, Y.B., Iwarsson, M., Otieno, D., Balkwill, K., Phillipson, P.B., Harley, R.M. & Willis, F. (2013). Flora Zambesiaca 8(8): 1-346. Royal Botanic Gardens, Kew.
  • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
  • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
  • Strugnell, A.M. (2006). A checklist of the Spermatophytes of Mt. Mulanje, Malawi Scripta Botanica Belgica 34: 1-199.
  • Suddee, S., Paton, A.J. & Parnell, J.A.N. (2005). Taxonomic Revision of the tribe Ocimeae Dumort (Lamiaceae) in continental South East Asia III. Ociminae Kew Bulletin 60: 3-75.
  • Velayos, M., Barberá, P., Cabezas, F.J., de la Estrella, M., Fero, M. & Aedo, C. (2014). Checklist of the vascular plants of Annobón (Equatorial Guinea) Phytotaxa 171: 1-78.
  • Welsh, S.L. (1998). Flora Societensis: 1-420. E.P.S. Inc. Utah.

Literature

Catálogo de Plantas y Líquenes de Colombia

  • ColPlantA (2021). "ColPlantA. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://www.colplanta.org/"

Flora of West Tropical Africa

  • Chev. Bot. 511 (as var. macrophyllum Briq.)
  • Morton in J. Linn. Soc. 58: 232 (1962).
  • Sp. Pl. 2: 1197 (1753)

Kew Backbone Distributions

  • (2020). file:///C:/Users/rg02kg/Downloads/7790-19423-1-PB.pdf epublication.
  • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Baksh-Comeau, Y., Maharaj, S.S., Adams, C.D., Harris, S.A., Filer, D.L. & Hawthorne, W.D. (2016). An annotated checklist of the vascular plants of Trinidad and Tobago with analysis of vegetation types and botanical 'hotspots' Phytotaxa 250: 1-431.
  • Berhaut, J. (1975). Flore illustrée du Sénégal 4: 1-625. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
  • Brundu, G. & Camarda, I. (2013). The Flora of Chad: a checklist and brief analysis PhytoKeys 23: 1-18.
  • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
  • Catarino, L., Martins, E.S., Diniz, M.A. & Pinto-Basto, M.F. (2006). Check-list da flora vascular do parque natural das Lagos de Cufada (Guiné-Bissau) Garcia de Orta, Série de Botânica 17: 97-141.
  • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
  • Fosberg, F.R. (1957). The Maldive islands, Indian Ocean Atoll Research Bulletin 58: 1-37.
  • Fosberg, F.R. (1975). Vascular plants of Aitutaki Atoll Research Bulletin 190: 73-84.
  • Harris, D.J. (2002). The vascular plants of the Dzanga-Sangha Reserve, Central African Republic: 1-274. National Botanic Garden (Belgium), Meise.
  • Jones, M. (1991). A checklist of Gambian plants: 1-33. Michael Jones, The Gambia College.
  • Lebrun, J.P., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Lorence, D.H. & Wagnwe, W.L. (2020). Flora of the Marquesas Islands 2: 413-1135. National Tropical Botanic Garden, Smithsonian, DRPF.
  • Mabberley, D.J. & De Kok, R.P.J. (2004). Labiatae Flore de la Nouvelle-Calédonie et Dépendances 25: 20-141. Muséum National d'Histoire Naturelle, Paris.
  • Morales, R. (2011). Les Labiadas (Lamiaceae) de Guinea Ecuatorial Anales del Jardin Botanico de Madrid 68: 199-223.
  • Pandey, R.P. & Dilwakar, P.G. (2008). An integrated check-list flora of Andaman and Nicobar islands, India Journal of Economic and Taxonomic Botany 32: 403-500.
  • Paton, A.J., Bramley, G., Ryding, O., Polhill, R.M., Harvey, Y.B., Iwarsson, M., Otieno, D., Balkwill, K., Phillipson, P.B., Harley, R.M. & Willis, F. (2013). Flora Zambesiaca 8(8): 1-346. Royal Botanic Gardens, Kew.
  • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
  • Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo, ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
  • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
  • Suddee, S., Paton, A.J. & Parnell, J.A.N. (2005). Taxonomic Revision of the tribe Ocimeae Dumort (Lamiaceae) in continental South East Asia III. Ociminae Kew Bulletin 60: 3-75.
  • Wagner, W.L., Herbst, D.R. & Sohmer, S.H. (1999). Manual of the Flowering Plants of Hawai'i, rev. ed., 1: 1-988. University of Hawai'i Press, Bishop Museum Press.
  • Welsh, S.L. (1998). Flora Societensis: 1-420. E.P.S. Inc. Utah.
  • Yuncker, T.G. (1959). Plants of Tonga Bernice P. Bishop Museum Bulletin 220: 1-283.

Flora of Somalia

  • Flora Somalia, (2000) Author: by O. Ryding [updated by M. Thulin 2008]
  • Paton in Kew Bull. 47: 403–435 (1992)

Flora of Tropical East Africa

  • F.W.T.A., ed. 2, 2: 452 (1963)
  • Fl. Rwanda 3: 324 (1985)
  • Holm & Hiltunen, Ocimum: 26 (1999).
  • K.B. 47: 411, fig. 1G, H (1992)
  • P.O.A. C: 350 (1895)
  • Sp. Pl.: 1197 (1753)
  • U.K.W.F. ed. 2: 296, t. 132 (1994)

Art and Illustrations in Digifolia
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Catálogo de Plantas y Líquenes de Colombia
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Colombian resources for Plants made Accessible
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Flora of Somalia
Flora of Somalia
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Flora of Tropical East Africa
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Flora of West Tropical Africa
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Herbarium Catalogue Specimens
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Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Bulletin
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Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

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