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This species is accepted, and its native range is Africa, India to Indo-China, Central & E. Jawa.
Orthosiphon marmoritis

[FTEA]

Lamiaceae (Labiatae), A.J. Paton, G. Bramley, O. Ryding, R.M. Polhill, Y.B. Harvey, M. Iwarsson, F. Willis, P.B. Phillipson, K. Balkwill, C.W. Lukhoba, D.F. Otieno, & R.M. Harley. Flora of Tropical East Africa. 2009

Type
Type: South India, Heyne s.n. (B†, holo.)
Morphology General Habit
Perennial herb, straggling, becoming somewhat shrubby, flowering from an early stage and developing a small rootstock with a number of ascending stems 0.2–1.5 m tall, not or hardly aromatic
Morphology Stem
Stems erect, quadrangular, normally well branched, with incipient shoots in many of the axils, retrorsely pubescent along the angles, more densely hairy above and occasionally with dense spreading indumentum overall
Morphology Leaves
Leaves petiolate, glandular-punctate; blade ovate to elliptic, mostly 1–4 × 0.6–3 cm, but shade forms up to 6–8 × 4–5 cm, crenate to serrate, acute or slightly acuminate to obtuse at apex, usually cuneate and shortly attenuate at base, occasionally more truncate, glabrous to pubescent above, pubescent especially along the veins beneath; petiole 5–10(–25) mm long
Morphology Reproductive morphology Inflorescences
Inflorescence soon lax, the 6-flowered verticils 8–14(–20) mm apart; bracts ovate-triangular, 2–3 mm long, erect above forming an inconspicuous apical coma, deflexed below; pedicels 1–2 mm long in flower, up to 4 mm in fruit
Morphology Reproductive morphology Flowers Calyx
Calyx conspicuously purple on upper side (posterior lip), green on underside, spreading at anthesis, 3–4 mm long, pubescent at base and along veins; fruiting calyx downward-pointing or vertically so, 5–7 mm long; posterior lip obtuse, minutely mucronate, shortly recurrent on the tube, 1.5–2(–3) mm long; lower lobes of the anterior lip shortly united, awn-like, 2–3 mm long, often 1 mm or so longer than lateral lobes
Morphology Reproductive morphology Flowers Corolla
Corolla usually pink, sometimes white or mauve, 8–10(–16) mm long; tube 6–8 mm long, straight, parallel-sided, dilating slightly towards throat; posterior lip perpendicular to tube; anterior lip cucullate, enclosing stamens, curving slightly upwards or horizontal, finally slightly deflexed and releasing stamens
Morphology Reproductive morphology Fruits
Nutlets brown, speckled darker, broadly ovoid, 0.9–1.2 mm long, producing a small amount of mucilage when wet.
Ecology
Wooded grassland, disturbed places, extending into bushland and more open dry forest; near sea-level to 2100 m
Conservation
Least concern; widely distributed
Note
Closely related to O. schimperi, see discussion there.
Distribution
Range: Tropical Africa and South Africa, Madagascar and tropical Asia from Sri Lanka and Peninsular India to China and Indonesia Flora districts: U1 U2 U3 U4 K1 K2 K3 K4 K5 K6 K7 T1 T2 T3 T5 T6 T7 T8

[KBu]

Suddee, S., A. J. Paton, & Parnell, J. (2005). Taxonomic Revision of Tribe Ocimeae Dumort. (Lamiaceae) in Continental South East Asia III. Ociminae. Kew Bulletin, 60(1), 3-75. Retrieved from http://www.jstor.org/stable/4110885

Distribution
Sri Lanka, Nepal, India, Burma?, China, Laos, Thailand?, Vietnam, Malaysia & Indonesia.
Ecology
In evergreen forest, on limestone rocks in forest; near sea level- 700 m. Flowering and fruiting all year round.
Morphology General Habit
Erect or ascending herb to 0.8 m tall
Morphology Leaves
Leaves ovate, papery, 20 - 90 x 10 - 50 mm, apex acute, base truncate or shortly cuneate, margin coarsely serrate, glabrous, puberulous or pubescent with sessile glands on both sides, glands and veins prominent beneath; petioles 8 - 45 mm long, pubescent
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens varying from included under anterior corolla lip to slightly exserted, anterior ones slightly longer
Morphology Reproductive morphology Flowers Calyx
Calyx 4 - 6 mm long at anthesis, 7 - 9 mm long in fruit; posterior lip orbicular or ovate-orbicular, apex rounded, acute, or apiculate, decurrent on tube, glabrous both sides; anterior lip with subulate teeth, lower teeth the same length or slightly longer than posterior, lateral slightly shorter than posterior; tube puberulous or pubescent outside, with or without gland-tipped hairs, sometimes glabrescent on upper side inside
Morphology Reproductive morphology Flowers Corolla
Corolla white or whitish-purple, 10 - 17 mm long, 3 times as long as calyx; posterior lip 3- lobed, midlobe emarginate; anterior lip slightly concave, ± equal in length to posterior; tube 7 - 12 mm long, dilated towards throat, straight or slightly curved, pubescent outside, hairs denser on anterior side
Morphology Reproductive morphology Flowers Disc
Disc with anterior side well- developed, apex acute, not exceeding ovary
Morphology Reproductive morphology Flowers Gynoecium Style
Style ± equal to the anterior stamens, clavate and cleft at apex
Morphology Reproductive morphology Fruits
Nutlets brown, ellipsoid, 1 - 1.2 mm long, minutely tuberculate or reticulate, producing mucillage when wet or not.
Morphology Reproductive morphology Inflorescences
Inflorescence terminal, up to 25 cm long, sometimes branched at base; axis puberulous or pubescent, with or without gland-tipped hairs; bracts sessile, subtending 2 - 3 flowers, forming a small apical coma, persistent, ovate, 2 - 3 mm long, shorter than pedicel in flower and fruit, acute, glabrous all over or adaxial glabrous, abaxial puberulous; pedicels 2.5 - 5 mm long at anthesis, 4 - 6 mm in fruit, puberulous, with or without gland-tipped hairs
Morphology Stem
Stems woody at base, simple or much branched, quadrangular or round-quadrangular, glabrous or glabrescent below, pubescent above
Note
Orthosiphon thymiflorus is highly variable in hairiness and stamen length. In Indian material, the leaves vary from glabrous to densely pubescent and the stamens are never exserted from the anterior corolla lip. In the Chinese and Indo-Chinese material, the stamens vary from being included in anterior corolla lip (type of O. sinensis Hemsl.) to slightly exserted, as in to the types of O. marmoritis (Plectranthus marmoritis Hance). However the leaves of Chinese and Indo-Chinese material are mostly thin and brittle, whether or not the stamens are exserted. Indian material can sometimes have similar leaves. There is therefore no clear discontinuity in leaves or in other vegetative characters. Doan (1936) used the exserted stamens as a means of separating Orthosiphon thymiflorus (O. tomentosus in his sense) from O. marmoritis but this cannot be upheld because there are intermediates between these two extremes. The formerly accepted name O. marmoritis is here reduced to synonymy. There was also confusion about the varietal names proposed by Sleesen (1959) under Orthosiphon thymiflorus for his revision of Malaysian Orthosiphon; var. tomentosus (Hook. f.) Sleesen should be read as var. tomentosus (Benth.) Sleesen and var. viscosus (Hook. f.) Sleesen should be read as var. viscosus (Benth.) Sleesen; the basionyms being O. tomentosus Benth. and O. viscosus Benth. respectively even though the type of O. tomentosus Benth. is in fact Endostemon viscosus.
Type
South India, Heyne s.n. (holotype B, destroyed).
Vernacular
Laotian: Pheng Deu Gieu, Phong Dieu Yeu, Nouat Meo, Ndi Kou (meo), Njo Tu Neng (meo). Vietnamese: Cay Tinh, Co Ham Huot, Cay Kinh Gioi Giai, Co Ham Huoi.

[FSOM]

M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS

Morphology General Habit
Herb with a woody base or subshrub, up to 0.2–1 m tall, slightly or densely pubescent
Morphology Leaves
Leaves petiolate; blade ovate, elliptic or oblong, up to 28–40 × 14–30 mm, apex rounded, obtuse or acute, margin crenate-dentate
Morphology Reproductive morphology Fruits Fruiting calyx
Fruiting calyx 5–7 mm long; upper lobe ± mucronate at apex; lower lobes 2–4 mm long
Morphology Reproductive morphology Flowers Corolla
Corolla 7–12 mm long, blue, violet, pink or rarely white; tube 6–7 mm long; lower lip 2–4 mm long
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens shorter than the lower lip of the corolla
Morphology Reproductive morphology Fruits
Nutlets c. 1.1 × 0.8 mm, producing mucilage when wet.
Distribution
N1, 2; widespread in tropical and southern Africa and tropical Asia.
Ecology
Altitude range 1000–2000 m.
Note
Orthosiphon suffrutescens is perhaps not distinct from O. schimperi and the Asiatic O. rubicundus (D. Don) Benth., and if this is the case, the species should be known under the latter name.

[FWTA]

Labiatae, J. K. Morton. Flora of West Tropical Africa 2. 1963

Morphology General Habit
A slender straggling subglabrous perennial herb
Ecology
Of deciduous and dry forest and thicket
Note
Frequently purple-tinged on the stem, leaves and inflorescence.

Doubtfully present in:

Myanmar, Thailand

Native to:

Angola, Benin, Botswana, Burundi, Central African Repu, China Southeast, Congo, Eritrea, Ethiopia, Ghana, India, Jawa, Kenya, KwaZulu-Natal, Laos, Madagascar, Malawi, Mozambique, Nigeria, Northern Provinces, Rwanda, Saudi Arabia, Somalia, Sri Lanka, Sudan, Swaziland, Tanzania, Togo, Uganda, Vietnam, Zambia, Zaïre, Zimbabwe

Orthosiphon thymiflorus (Roth) Sleesen appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Dec 1, 2012 Hemp [5642], Tanzania K000735701
Jan 1, 2009 Collenette [7153] K000248478
Nov 1, 2005 Schweinfurth, G. [3818] K000347188
Nov 1, 2005 Schweinfurth, G. [2765], Sudan K000347189
Nov 1, 2005 Holst, C. [3561a], Tanzania K000347200
Nov 1, 2005 Monteiro [s.n.], Angola K000347209 Unknown type material
Nov 1, 2005 Hildebrandt, J.M. [2747], Kenya K000347197 Unknown type material
Nov 1, 2005 Lort Phillips [306], Somalia K000347190 Unknown type material
Nov 1, 2005 Wakefield, T. [s.n.], Kenya K000347199 Unknown type material
Nov 1, 2005 Hildebrandt, J.M. [2746], Kenya K000347198 Unknown type material
Apr 1, 1964 Evans, I.H.N. [s.n.], Malaysia K000897133
Apr 1, 1964 Ridley, H.N. [s.n.], Malaysia K000897132
Hildebrandt, J.M. [1428], Somalia K000347191
Wallich, N. [s.n.], India K000911634 syntype
s.coll. [2723] K000911636
s.coll. [s.n.] K000911637
Clarke, C. B. [10461 B] K000911639
Gossweiler, J. [8375], Angola K001008854
[Youlkes, G.] [s.n.] K000911635 Unknown type material
Wallich, N. [s.n.], India K000911633 syntype
Wallich, N. [s.n.] K000911649 syntype
Wight [2107], India K000911638
Wigh [2501] K000911648 holotype
Wallich, N. [s.n.] K001080747
Faulkner, H.G. [A.21], Angola K001008856
Gossweiler, J. [9217], Angola K001008855
Faulkner, H.G. [A.87], Angola K001008853
s.coll. [Cat. no. 2724], India Orthosiphon glabratus K001116951
s.coll. [Cat. no. 2724] Orthosiphon glabratus K001116953
s.coll. [Cat. no. 2724], India Orthosiphon glabratus K001116952
Jan 1, 1989 Bolus, H. [11011], South Africa Orthosiphon suffrutescens K000347223 Unknown type material
Jan 1, 1989 Davy, B. [1744], South Africa Orthosiphon suffrutescens K000347227
Jan 1, 1989 Wilms, F. [2189], South Africa Orthosiphon suffrutescens K000347224
Medley Wood, J. [4963], South Africa Orthosiphon suffrutescens K000347225 Unknown type material
Wilms, F. [1113], South Africa Orthosiphon suffrutescens K000347226 Unknown type material
Jan 1, 1999 s.coll. [s.n.], Indonesia Orthosiphon petiolaris K000897110 isotype
Jan 1, 1999 s.coll. [s.n.], Indonesia Orthosiphon petiolaris K000897109 isotype
Jan 1, 2011 Nguyen [HNK-1417(de Kok 1217)], Vietnam Orthosiphon marmoritis K000249939
Ford, C. [146], China Orthosiphon marmoritis K000911671 holotype
Sampson, T. [17725], China Orthosiphon marmoritis K000911672 isotype
s.coll. [Cat. no. 2723], India Orthosiphon viscosus K001116950
s.coll. [Cat. no. 2723] Orthosiphon viscosus K001116949
Chancellor, R.J. [157], Uganda Orthosiphon australis 2767.000
Drummond, R.B. [2881], Tanzania Orthosiphon australis 4781.000

First published in Reinwardtia 5: 42 (1959)

Accepted by

  • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
  • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
  • Hedberg, I., Kelbessa, E., Edwards, S., Demissew, S. & Persson, E. (eds.) (2006). Flora of Ethiopia and Eritrea 5: 1-690. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.
  • Paton, A.J., Bramley, G., Ryding, O., Polhill, R., Harvey, Y., Iwarsson, M., Willis, F., Phillipson, P., Balkwill, K., Lukhoba, C., Otiend, D & Harley (2009). Lamiaceae (Labiatae) Flora of Tropical East Africa: 1-430.
  • Suddee, S., Paton, A.J. & Parnell, J.A.N. (2005). Taxonomic Revision of the tribe Ocimeae Dumort (Lamiaceae) in continental South East Asia III. Ociminae Kew Bulletin 60: 3-75.

Literature

Kew Backbone Distributions

  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Bramley, G.L.C. (2019). Flora Malesiana 23: 1-444. Noordhoff-Kolff N.V., Djakarta.
  • Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.
  • Suddee, S., Paton, A.J. & Parnell, J.A.N. (2005). Taxonomic Revision of the tribe Ocimeae Dumort (Lamiaceae) in continental South East Asia III. Ociminae Kew Bulletin 60: 3-75.

Flora of Tropical East Africa

  • Fl. Eth. 5: 582 (2006).
  • Fl. Madag. 175: 128 (1998)
  • K.B. 60: 7 (2005)
  • Reinwardtia 5: 42 (1952)

Flora of Somalia
Flora of Somalia
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Flora of Tropical East Africa
Flora of Tropical East Africa
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Flora of West Tropical Africa
Flora of West Tropical Africa
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Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Bulletin
Kew Bulletin
http://creativecommons.org/licenses/by-nc-sa/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Science Photographs
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