Crossandra Salisb.

First published in Parad. Lond. 1: t. 12 (1805)
This genus is accepted
The native range of this genus is Tropical Africa to Sri Lanka.

Descriptions

Acanthaceae (part 1), Kaj Vollesen, Flora of Tropical East Africa, 2008

Morphology General Habit
Perennial (rarely annual) herbs, subshrubs or shrubs.
Morphology Leaves
Leaves opposite or in whorls of 4, from below middle usually gradually narrowed into a long-decurrent base.
Morphology Reproductive morphology Flowers
Flowers single, in dense sessile or pedunculate, terminal or axillary spikes; bracts imbricate, in 4 rows, persistent, papery to coriaceous, with straight or recurved often pungent tip or 3-lobed at apex, margin entire, toothed, spinulose or setose; lower sterile and smaller; bracteoles 2, usually linear to lanceolate and acuminate to cuspidate, usually herbaceous.
Morphology Reproductive morphology Flowers Calyx
Calyx glumaceous, divided to base in 5 unequal sepals which are thickened and horny at base; dorsal sepal 2-veined and 2-toothed (in C. cephalostachya and leucodonta 1-veined and 1-toothed), broader than and often enveloping the rest, ventral and lateral 1-veined and 1-toothed, lateral smaller than ventral.
Morphology Reproductive morphology Flowers Corolla
Corolla puberulous outside; tube linear, usually widened above stamens, with a band of hairs at insertion of stamens; limb and top of tube split posteriorly to give a single 5-lobed lower lip, lobes oblong, obtuse, middle broadest and often emarginate.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 4, included, usually inserted ± 1/3 down the tube, subsessile; anthers 1-thecous, oblong, usually rounded to finely apiculate, finely bearded.
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary 2-locular with 2 ovules per locule, glabrous or minutely hairy near top; style usually glabrous, gibbous or not below the obliquely trumpet-shaped or obscurely 2-lobed stigma which is at the same height as anthers.
Morphology Reproductive morphology Fruits
Capsule 4-seeded, ellipsoid, sessile, glumaceous, glabrous and shiny.
Morphology Reproductive morphology Seeds
Seed discoid, obliquely ovoid, covered by large appressed laciniate scales.
Distribution
Range: 52 species in tropical Africa and Madagascar with 1 species extending to India and 1 species endemic in Arabia.
[FTEA]

M. Thulin et al. Flora of Somalia, Vol. 1-4 [updated 2008] https://plants.jstor.org/collection/FLOS

Morphology Reproductive morphology Flowers Corolla
Corolla white, yellow, orange or red, limb and top of tube split on the back and forming a single 5-lobed lower lip
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 4, included; anthers 1-celled, finely bearded
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary 2-celled with 2 ovules per cell; style with obliquely trumpet-shaped stigma
Morphology Reproductive morphology Fruits
Capsule ellipsoid, 4-seeded
Morphology Reproductive morphology Seeds
Seeds flattened, covered by fringed scales and sticky when wet.
Distribution
Some 50 species in tropical Africa, Madagascar, south-western part of the Arabian Peninsula, and in southern Asia.
Morphology General Habit
Herbs, subshrubs or shrubs
Morphology Leaves
Leaves opposite or seemingly in whorls of 4, usually narrowed from below middle into a long attenuate base
Morphology Reproductive morphology Inflorescences
Flowers in dense terminal or axillary spikes; bracts in 4 rows, persistent, often with pungent tip, the margin entire, toothed, spinulose or setose, the lower bracts sterile, smaller than the fertile ones; bracteoles 2, narrow
Morphology Reproductive morphology Flowers Calyx
Calyx deeply 5-lobed; lobes unequal, the upper 2-toothed, broader than the others and often enveloping them
[FSOM]

J. R. Timberlake, E. S. Martins (2013). Flora Zambesiaca, Vol 8 (5) Acanthaceae Part 1. Royal Botanic Gardens, Kew

Morphology General Habit
Perennial herbs, subshrubs or shrubs
Morphology Leaves
Leaves opposite or in pseudo-whorls of 4, often gradually narrowed from below middle into a long-decurrent base, usually entire
Morphology Reproductive morphology Flowers
Flowers single, in dense sessile or pedunculate, terminal or axillary spiciform cymes
Morphology Reproductive morphology Inflorescences Bracts
Bracts imbricate, in 4 rows, persistent, papery to coriaceous, with a straight or recurved often pungent tip or 3-lobed at apex, margin entire, toothed or spinulose; lowermost sterile and smaller, often distant
Morphology Reproductive morphology Inflorescences Bracteoles
Bracteoles 2, usually linear to lanceolate and acuminate to cuspidate, usually herbaceous
Morphology Reproductive morphology Flowers Calyx
Calyx glumaceous, divided to base in 5 unequal sepals, thickened and horny at base; dorsal sepal 2-veined and 2-toothed, broader than and often enveloping the rest, ventral and lateral 1-veined and 1-toothed, lateral usually smaller than ventral-Corolla white, yellow, orange or light to bright red, puberulous outside; tube linear, widened above stamens and with band of hairs at stamen insertion; limb and top of tube split dorsally to give a single 5-lobed lower lip; lobes oblong, obtuse, middle broadest and often emarginate
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 4, usually inserted ± 1/3 way down tube, subsessile; anthers included, 1-thecous, oblong, usually rounded to finely apiculate, finely bearded
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary 2-locular with 2 ovules per locule, glabrous or minutely hairy near top; style usually glabrous, gibbous or not below the obliquely trumpet-mouthed or obscurely 2-lobed stigma, which is same height as anthers
Morphology Reproductive morphology Fruits
Capsule 4-seeded, ellipsoid, sessile, glumaceous, glabrous and shiny
Morphology Reproductive morphology Seeds
Seed discoid, obliquely ovoid, covered by large appressed laciniate scales.
Distribution
A genus of 52 species, mainly in tropical Africa and Madagascar with one species extending to India and one endemic to Arabia.
[FZ]

Sources

  • Art and Illustrations in Digifolia

    • Digital Image © Board of Trustees, RBG Kew
  • Flora Zambesiaca

    • Flora Zambesiaca
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Flora of Somalia

    • Flora of Somalia
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Flora of Tropical East Africa

    • Flora of Tropical East Africa
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Herbarium Catalogue Specimens

    • 'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
  • Kew Backbone Distributions

    • The International Plant Names Index and World Checklist of Vascular Plants 2025. Published on the Internet at http://www.ipni.org and https://powo.science.kew.org/
    • © Copyright 2023 World Checklist of Vascular Plants. http://creativecommons.org/licenses/by/3.0
  • Kew Names and Taxonomic Backbone

    • The International Plant Names Index and World Checklist of Vascular Plants 2025. Published on the Internet at http://www.ipni.org and https://powo.science.kew.org/
    • © Copyright 2023 International Plant Names Index and World Checklist of Vascular Plants. http://creativecommons.org/licenses/by/3.0