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This species is accepted, and its native range is Kenya to Mozambique, W. Indian Ocean.
Hymenaea verrucosa

[ILDIS]

International Legume Database and Information Service

Conservation
Not Threatened
Ecology
Africa: Zanzibar-Inhambane regional transition zone; forest, Zanzibar-Inhambane Regional transition zone; Transition woodland.
Morphology General Habit
Perennial, Not climbing, Tree
Vernacular
Analomanta, Copalier, Gum Copal, Mandrirofo, Mandrofo, Mandrorofo, Tandrofo, Tandroroho, Zanzibar Copal

[FZ]

Leguminosae, R.K. Brummitt, A.C. Chikuni, J.M. Lock and R.M. Polhill. Flora Zambesiaca 3:2. 2007

Morphology General Habit
Large tree (or occasionally a shrub) to at least 20 m, with broad crown.
Morphology Branches
Young branchlets pale brown, lenticellate, glabrous.
Morphology Leaves
Leaves unijugate; stipules tiny, triangular with ciliate margins, early caducous; petiole 1.2–3 cm long, glabrous; leaflets 5.3–10.5 × 2.8–5.5 cm, ovate-elliptic, asymmetric, the inner margin weakly curved, cuneate at the base, the outer strongly curved and cuneate to subcordate at the base, apex bluntly acuminate, glabrous; midrib distinct, venation prominulous on both surfaces, resin vesicles (dark spots) scattered in the areolae beneath.
Morphology Reproductive morphology Inflorescences
Inflorescences axillary and terminal, paniculate; axis terete, densely to sparsely appressed-pubescent, glabrescent; bracts and bracteoles c.2.5 mm in diameter, subcircular, concave, very early caducous.
Morphology Reproductive morphology Flowers
Flowers c.15 mm long; pedicels 5–7 mm long, terete.
Morphology Reproductive morphology Flowers Calyx
Calyx lobes 4, c.10 × 5–7 mm, broadly elliptic to obovate, densely pubescent outside, densely sericeous inside.
Morphology Reproductive morphology Flowers Corolla
Petals 5; the three upper ones white, clawed, the limb c.9 × 8 mm, suborbicular, the claw c.10 mm long; the two lower ones minute, c.1.5 mm long, oblong.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 10; filaments 12–14 mm long, glabrous; anthers dorsifixed.
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary 4-ovulate, oblong, c.3 × 2 mm, pilose; stipe very short, densely spreading-hairy at the apex, glabrous below; style c.6 mm long, glabrous, stigma capitate.
Morphology Reproductive morphology Fruits
Pod c.5 × 3 cm, ovoid to ellipsoidal, glabrous except for a ring of hairs at the extreme base, with numerous superficial resin-filled vesicles, 1–3-seeded, indehiscent.
Morphology Reproductive morphology Seeds
Seed dark brown, shiny, c.20 × 12 mm, ellipsoidal, enclosed in the cream-coloured mealy endocarp.

[FTEA]

Leguminosae, J. B. Gillett, R. M. Polhill & B. Verdcourt. Flora of Tropical East Africa. 1971

Morphology General Habit
Tree 6–24 m. high (–40 m., fide Léonard, l.c., 1950); bark grey or greyish-white, smooth or rough; young branchlets puberulous to glabrous.
Morphology Leaves
Leaflets ovate to elliptic or oblong-elliptic, asymmetric, 3.5–12 cm. long, 2–5.7 cm. wide (smaller ones often present with the inflorescence, larger (to 15 × 7.4 cm.) on juvenile shoots), coriaceous, glabrous or subglabrous, obtusely pointed to shortly subacuminate at apex.
Morphology Reproductive morphology Inflorescences
Panicle up to 35 × 25 cm., but often much smaller; ultimate branches and pedicels densely and shortly whitish-pubescent.
Morphology Reproductive morphology Flowers
Flowers white.
Morphology Reproductive morphology Flowers Calyx
Sepals 7–11 mm. long, densely appressed-pubescent outside and silvery-silky inside.
Morphology Reproductive morphology Flowers Corolla
Larger petals ± 1.5–2 cm. long; lamina ± 8–10 mm. in diameter.
Morphology Reproductive morphology Fruits
Pods ovoid-oblong, ellipsoid-oblong, or obovoid, 2.5–5 cm. long, 1.5–3 cm. wide.
Morphology Reproductive morphology Seeds
Seeds 1.3–1.8 × 0.9–1.2 cm.
Figures
Fig. 25, p. 133.
Habitat
On or near the coast, in lowland dry evergreen forest, woodland and coastal evergreen bushland; 15–240 m.
Distribution
K7 T3 T6 T8 Z

[ILDIS]
Use
Chemical products, Domestic, Food and Drink, Medicine, Wood

Native to:

Comoros, Kenya, Madagascar, Mauritius, Mozambique, Réunion, Tanzania

Introduced into:

Benin, Borneo, Ghana, India, Jawa, Senegal, Seychelles, Sri Lanka, Taiwan

Hymenaea verrucosa Gaertn. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Jan 1, 1997 Capuron, R. [8934-SF], Madagascar K000297192
Mar 28, 1989 Groenendijk, E.M.C. [2159], Mozambique K000297191
Jan 1, 1985 Cadet, Th. [6630], Reunion K000297190
Dupuy, D.J. [M 203], Madagascar 63590.000
Peters [s.n.], Mozambique K000417811
Du Puy, D.J. [M203], Madagascar K000297189
Du Puy, D.J. [M13], Madagascar K000297188

First published in Fruct. Sem. Pl. 2: 306 (1791)

Accepted by

  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Barthelat, F. (2019). La flore illustrée de Mayotte: 1-687. Biotope éditions.
  • Brummitt, R.K., Chikuni, A.C., Lock, J.M. & Polhill, R.M. (2007). 61. Leguminosae Subfamily Caesalpinioideae Flora Zambesiaca 3(2): 1-218. Royal Botanic Gardens, Kew.
  • Du Puy, D.J., Labat, N.-N., Rabevohitra, R., Villiers, J.-F., Bosser, J. & Moat, J. (2002). The Leguminosae of Madagascar: 1-737. Royal Botanic Gardens, Kew.
  • Kumar, S. & Sane, P.V. (2003). Legumes of South Asia. A Checklist: 1-536. Royal Botanic Gardens, Kew.
  • Lock, J.M. & Ford, C.S. (2004). Legumes of Malesia a Check-List: 1-295. Royal Botanic Gardens, Kew.
  • Lock, J.M. (1989). Legumes of Africa a check-list: 1-619. Royal Botanic Gardens, Kew.
  • Wu, Z. & Raven, P.H. (eds.) (2010). Flora of China 10: 1-642. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).

Literature

Kew Backbone Distributions

  • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • Barthelat, F. (2019). La flore illustrée de Mayotte: 1-687. Biotope éditions.
  • Berhaut, J. (1975). Flore illustrée du Sénégal 4: 1-625. Gouvernement du Sénégal, Ministère du développement rural direction des eaux et forêta, Dakar.
  • Brummitt, R.K., Chikuni, A.C., Lock, J.M. & Polhill, R.M. (2007). 61. Leguminosae Subfamily Caesalpinioideae Flora Zambesiaca 3(2): 1-218. Royal Botanic Gardens, Kew.
  • Castle, G.E. (1994). Flore des Seychelles Dicotylédones: 1-663. Orstom Editions.
  • Kumar, S. & Sane, P.V. (2003). Legumes of South Asia. A Checklist: 1-536. Royal Botanic Gardens, Kew.
  • Lock, J.M. & Ford, C.S. (2004). Legumes of Malesia a Check-List: 1-295. Royal Botanic Gardens, Kew.
  • Lock, J.M. (1989). Legumes of Africa a check-list: 1-619. Royal Botanic Gardens, Kew.
  • Wu, Z. & Raven, P.H. (eds.) (2010). Flora of China 10: 1-642. Science Press (Beijing) & Missouri Botanical Garden Press (St. Louis).

International Legume Database and Information Service

  • Backer, C. A. (1963). Flora of Java. Groningen, The Netherlands
  • Brenan, J. P. M. (1959). Mimosoideae. In: Fl. Trop. East Africa. Hubbard & Milne-Redhead
  • Brenan, J. P. M. (1967). Caesalp. In: Flora Trop. East Africa. Milne-Redhead & Polhill
  • Dale, I. R. & Greenway, P. J. (1961). Kenya Trees and Shrubs
  • Ding Hou, Larsen, S & Larsen, K (1996). Flora Malesiana I, 12, 2:409-784. Caesalpiniaceae.
  • Du Puy, D. J. et al. (2002). The Leguminosae of Madagascar. RBG Kew.
  • Polhill, R, M. (1990). Legumineuses. In: Flore des Mascareignes, Vol 80. J. Bosser et a
  • Robertson, S. A. (1989). Flowering Plants of Seychelles. Kew
  • Rudd, V. E. (1991). Flora of Ceylon 7: 34-107. Caesalpinioideae
  • White, F. (1983). The Vegetation of Africa
  • de Cordemoy, E. J. (1895). Flore de l'ile de la Reunion

Flora Zambesiaca
Flora Zambesiaca
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Flora of Tropical East Africa
Flora of Tropical East Africa
http://creativecommons.org/licenses/by-nc-sa/3.0

Herbarium Catalogue Specimens
Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

International Legume Database and Information Service
International Legume Database and Information Service (ILDIS) V10.39 Nov 2011
http://creativecommons.org/licenses/by/3.0

Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Science Photographs
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