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This species is accepted, and its native range is Angola.

[KBu]

Darbyshire, I., Tripp, E.A. & Chase, F.M. (2019). A taxonomic revision of Acanthaceae tribe Barlerieae in Angola and Namibia. Part 1. Kew Bulletin 74: 5. https://doi.org/10.1007/s12225-018-9791-0

Conservation
Barleria namba was first discovered in 2016 and is known only from the type locality of Mt Namba. This mountain has remnant patches of Afromontane forest but some areas have been cleared and are seasonally burned and subject to cattle grazing. The steeper rocky slopes on which this species is found are better protected from some of these impacts and it is likely that these rocky areas support a more open vegetation even in the absence of human disturbance. That said, in view of its extremely small range and the plausible threats from burning and cattle grazing, this species is assessed as Vulnerable — VU D2.
Distribution
Endemic to western Angola (Cuanza Sul Prov.), known only from Mt Namba.
Ecology
Barleria namba is recorded from open rocky hillslopes subject to grazing, at ± 2000 m elevation.
Morphology General Habit
Spreading subshrub, much branched, to 60 cm tall; stems reddish-brown, densely pale-buff pubescent with mixed long ascending bulbous-based hairs and shorter more spreading hairs, both most dense on two opposite sides
Morphology Leaves
Leaves sessile or petiole to 4 mm long; blade (?only seen in immature state) elliptic or somewhat ovate, 2.7 – 4 × 1.4 – 1.9 cm (l:w ratio 1.8 – 2.1:1), base cuneate or obtuse, margin entire, apex acute, surfaces with numerous long pale buff hairs throughout and with interspersed shorter fine hairs beneath; lateral veins 4 pairs, prominent beneath, fine reticulate venation also conspicuous beneath, dark in dry state
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens inserted 6 – 6.5 mm from base of corolla tube; filaments 27 – 30 mm long; anthers exserted, 3.6 – 4 mm long; lateral staminodes 3 – 3.5 mm long, glabrous except for minute subsessile glands towards base, antherodes 0.7 – 1 mm long
Morphology Reproductive morphology Flowers Calyx
Calyx green or purplish with conspicuous darker subparallel primary venation; anterior lobe elliptic or elliptic-lanceolate, 15 – 19 × 5 – 6.5 mm in flower, up to 22.5 mm long in young fruit, base cuneate, margin entire, somewhat involute, apex acute-mucronate or unevenly notched, external surface with dense short viscid-glandular hairs and with interspersed eglandular hairs, these longest along the main veins and margin; posterior lobe as anterior lobe but more lanceolate, 18.5 – 21 × 6 – 7 mm in flower, up to 24.5 mm long in young fruit; lateral lobes linear-lanceolate, 13 – 15 mm long
Morphology Reproductive morphology Flowers Corolla
Corolla white tinged pale pink-purple and with darker markings in throat, drying dark blue with darker venation, 39 – 41 mm long, glandular-pilose and eglandular-pubescent externally; tube 21 – 23 mm long, cylindrical in basal portion where 3 – 4 mm wide, campanulate above insertion point of stamens, mouth 7.5 – 8.5 mm wide; limb in marked “2+3” configuration; abaxial lobe barely offset, broadly obovate, 16.5 – 17 × 12.5 – 13 mm, apex shallowly emarginate, lateral lobes elliptic or obovate-elliptic, 17 – 18.5 × 9 – 11 mm, apices obtuse or rounded, adaxial lobes lanceolate 15.5 – 17 × 3.8 – 5 mm, pair fused in basal 4.5 – 5 mm, apices markedly acute or attenuate
Morphology Reproductive morphology Flowers Gynoecium Ovary
Ovary with few pale straight hairs at apex; style glabrous; stigma subcapitate but slender, 0.25 – 0.3 mm long
Morphology Reproductive morphology Fruits
Capsule only seen in immature state, apparently 4-seeded, glabrous except for a few minute hairs at apex.
Morphology Reproductive morphology Inflorescences
Inflorescences of crowded terminal spikes on short lateral shoots, each comprising a series of sessile single-flowered cymes; bracts foliaceous but much reduced, 7 – 16.5 × 3 – 8.5 mm, apex spinulose, indumentum as leaves but with additional short glandular hairs; bracteoles linear-lanceolate, 11 – 14.5 × 1.5 – 2 mm, margin entire, apex spinulose, surface conduplicate, green with pale and prominent midrib, indumentum as bracts, glandular hairs numerous
Note
This species is most similar to Barleria rupicola and B. crassa but is easily separated from them by (i) the corolla limb being markedly zygomorphic with lanceolate adaxial lobes fused in the lower third (vs corolla limb only weakly zygomorphic, adaxial lobes elliptic, not clearly fused at base); (ii) the inflorescence being viscid-glandular throughout (vs glandular hairs absent or sparse and not viscid); (iii) having well-defined spiciform inflorescences with the bracts markedly reduced in comparison to the leaves (inflorescences can be crowded towards stem apices in B. rupicola and B. crassa but the bracts are foliaceous, not reduced); and (iv) having a capitate stigma to 0.3 mm long (vs linear or clavate, 0.8 – 2 mm long). The species epithet “namba” refers to the type locality of Mt Namba. This striking new species appears to be allied to Barleria rupicola and B. crassa as it shares with these species a funnel-shaped throat to the corolla that is longer than the cylindrical base, a “2+3” corolla configuration, linear-lanceolate, conduplicate bracteoles that lack marginal spines or teeth, and straight hairs at the insertion point of the style, as well as having a similar ecology. However, it is easily separated from these species by the characters listed in the Recognition section. The corolla form approaches that of B. solitaria (sp. 20 below) although it lacks the purple speckling on the adaxial lobes which are fused for a shorter distance and it has a broader funnel-shaped tube.
Type
Angola, Cuanza Sul Prov., Mt Namba, c. 34 km W of Cassongue, N face of mountain above Kanhala village, fl. 12 June 2016, Goyder, Maiato & Luís 8660 (holotype K! [K001295764]; isotype LUBA).

Native to:

Angola

Barleria namba I.Darbysh. appears in other Kew resources:

First published in Kew Bull. 74(1)-5: 53 (2019)

Accepted by

  • Govaerts, R., Nic Lughadha, E., Black, N., Turner, R. & Paton, A. (2021). The World Checklist of Vascular Plants, a continuously updated resource for exploring global plant diversity. https://doi.org/10.1038/s41597-021-00997-6 Scientific Data 8: 215.

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Kew Backbone Distributions
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

Kew Bulletin
Kew Bulletin
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Kew Names and Taxonomic Backbone
The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
© Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0